Research Article | | Peer-Reviewed

Gathering Pearls from the Lethe: Nomenclatural Novelties for Nothotaxa

Published in Plant (Volume 14, Issue 1)
Received: 2 January 2026     Accepted: 14 January 2026     Published: 27 January 2026
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Abstract

The integration of fine-scale taxonomic results into a comprehensive phylogenetic classification remains an ongoing challenge in modern systematics, particularly for infraspecific variants and interspecific hybrids, which are often overlooked. To address the resulting nomenclatural gaps and instabilities, this study provides a significant update by introducing numerous nomenclatural novelties for selected vascular plant genera, with a special focus on nothotaxa. In total, this work validates 452 nomenclatural novelties across 31 families, including Aizoaceae, Amaranthaceae, Amaryllidaceae, Apiaceae, Apocynaceae, Asteraceae, Berberidaceae, Cactaceae, Cistaceae, Crassulaceae, Cupressaceae, Cyperaceae, Equisetaceae, Fabaceae, Gesneriaceae, Melanthiaceae, Orchidaceae, Poaceae, Ranunculaceae and Saxifragaceae, ect. Among them are the establishment of 12 new nothogenera: × Osteolaena [Amaranthaceae], × Chrysania, × Cicuca, × Ozodes and × Psepaurea [Asteraceae], × Turdenicarpus [Cactaceae], × Astoxytropis [Fabaceae], × Eucongia [Gesneriaceae], × Pseudillium [Melanthiaceae], × Batepalum [Orchidaceae], × Rankwithia [Ranunculaceae] and × Mitarella [Saxifragaceae]; 59 nothoranks [five nothosubgenera, 42 nothosections, six nothosubsections and six nothoseries] under seven nothogenera [× Anacamptorchis, × Dactylocamptis, × Dactylodenia, × Gymnanacamptis, × Orchidactylorhiza, × Orchigymnadenia and × Orchimantoglossum] for the first time and as additions to five natural genera [Cattleya, Dactylorhiza, Paphiopedilum, Phalaenopsis and Pulsatilla], respectively; moreover, 360 new nomenclatural acts are proposed, covering 139 new combinations, 209 new combinaton and status, six new status, and six new replacement names; besides, nine names are synonymized under eight taxa. These changes reconcile hybrid taxonomy with contemporary phylogenetic insights, stabilize nomenclature and provide a crucial reference to support accurate identification, biodiversity documentation, and conservation efforts.

Published in Plant (Volume 14, Issue 1)
DOI 10.11648/j.plant.20261401.11
Page(s) 1-23
Creative Commons

This is an Open Access article, distributed under the terms of the Creative Commons Attribution 4.0 International License (http://creativecommons.org/licenses/by/4.0/), which permits unrestricted use, distribution and reproduction in any medium or format, provided the original work is properly cited.

Copyright

Copyright © The Author(s), 2026. Published by Science Publishing Group

Keywords

New Combinations, New Status, Replacement Name, Nothotaxa, Rank

1. Introduction
Building upon the broadly established phylogenetic framework at higher taxonomic levels, the continuous publication of new studies at the genus and species level necessitates the integration of their results into a complete classification down to the lowest ranks. This process creates a demand for new nomenclatural combinations, especially for infraspecific variations (subspecies, varieties, forms) and hybrids, which are typically overlooked in molecular phylogenetic studies. These often poorly differentiated microtaxa can possess significant local heritage value that risks being lost when they are subsumed into broader taxa. Furthermore, updating major nomenclatural databases requires incorporating findings from these numerous publications. Despite phylogenetic classification becoming standard in modern biology, the practical integration of fine-scale nomenclatural results remains an incomplete and challenging task, highlighting the need to validate new names for these overlooked ranks to support future identification and conservation.
The establishment of a stable nomenclature is particularly critical and challenging for nothospecies (interspecific hybrids), given their significant practical importance. Their value spans the creation of ornamental horticultural plants, the development of improved food crops, and their potential as medical resources. However, their nomenclature is frequently destabilized by ongoing taxonomic revisions driven by advances in molecular systematics, which routinely redefine genera through lumping and splitting. This instability underscores an urgent need for comprehensive compilations that reconcile hybrid taxonomy with contemporary phylogenetic classifications. Systematic efforts to document nothogeneric names and establish new nomenclatural arrangements are therefore essential to provide a reliable reference framework and accommodate newly described hybrids.
The taxonomical/nomenclatural concept of the current checklist follows the following principles:
Family-level. The author accepted the family circumscription from the classification systems APG IV for angiosperm, PPG I for lycophytes and fern allies, and a revised classification scheme for gymnosperm.
Genus-level. It is widely known, that one of the most difficult decisions concerns the choice of generic concepts, which is the main source of name changes . To avoid conflicts between contrasting points of view, the author applied the generic concept following the POWO.
Species-level. The acceptance of a genus is contingent on the specific database referenced. In contrast, defining a species often relies heavily on subjective expert knowledge. An exception exists when a species has undergone definitive phylogenetic analysis; its taxonomic status becomes stable once confirmed by authoritative molecular, morphological, or other studies. Entities involed here follow POWO and recent literature.
Nomenclature-level. Madrid Code of ICN has been followed in making necessary nomenclatural decisions. Unless more recent information is available, author attributions of scientific plant names cited follows IPNI, albeit in the notation required by the journal. To achieve precise writing, only selected references are provided for crucial taxonomic information.
2. Nomenclatural Treatment
2.1. Nomenclatural Novelties at Generic Level
Amaranthaceae
Osteolaena Z.H.Feng, nothogen. nov. — Parentage: Osteocarpum × Sclerolaena.
Asteraceae
× Chrysania Z.H.Feng, nothogen. nov. — Parentage: Ajania × Chrysanthemum.
× Cicuca Z.H.Feng, nothogen. nov. — Parentage: Cicerbita × Lactuca.
× Ozodes Z.H.Feng, nothogen. nov. — Parentage: Anaphalioides × Ozothamnus.
× Psepaurea Z.H.Feng, nothogen. nov. — Parentage: Centaurea × Psephellus.
Cactaceae
× Turdenicarpus Z.H.Feng, nothogen. nov. — Parentage: Kadenicarpus × Turbinicarpus.
Fabaceae
× Astoxytropis Z.H.Feng & K.W.Jiang, nothogen. nov. — Parentage: Astragalus × Oxytropis.
Gesneriaceae
× Eucongia Z.H.Feng, nothogen. nov. — Parentage: Eucodonia × Sinningia.
Melanthiaceae
× Pseudillium Z.H.Feng, nothogen. nov. — Parentage: Pseudotrillium × Trillium.
Orchidaceae
× Batepalum J.M.H.Shaw & Z.H.Feng, nothogen. nov. — Parentage: Batemannia × Zygosepalum.
Ranunculaceae
Rankwithia Z.H.Feng, nothogen. nov. — Parentage: Beckwithia × Ranunculus.
Saxifragaceae
× Mitarella Z.H.Feng, nothogen. nov. — Parentage: Mitella × Tiarella.
2.2. Nomenclatural Novelties at Infrageneric Level
Orchidiaceae
× Anacamptorchis E.G.Camus, J. Bot. (Morot) 6: 113 (1892).
× Anacamptorchis nothosect. Andranorchis Z.H.Feng, nothosect. nov. — Parentage: Anacamptis sect. Andranthus × Orchis sect. Orchis.
× Anacamptorchis nothosect. Coriorchis Z.H.Feng, nothosect. nov. — Parentage: Anacamptis sect. Coriophorae × Orchis sect. Orchis.
× Anacamptorchis nothosect. Masanthus Z.H.Feng, nothosect. nov. — Parentage: Anacamptis sect. Phalaenanthus × Orchis sect. Masculae.
× Anacamptorchis nothosect. Moriculae Z.H.Feng, nothosect. nov. — Parentage: Anacamptis sect. Morianthus × Orchis sect. Masculae.
× Anacamptorchis nothosect. Moriorchis Z.H.Feng, nothosect. nov. — Parentage: Anacamptis sect. Morianthus × Orchis sect. Orchis.
× Anacamptorchis nothosect. Morivinciales Z.H.Feng, nothosect. nov. — Parentage: Anacamptis sect. Morianthus × Orchis sect. Provinciales.
× Anacamptorchis nothosect. Phalaenorchis Z.H.Feng, nothosect. nov. — Parentage: Anacamptis sect. Phalaenanthus × Orchis sect. Orchis.
× Anacamptorchis nothosect. Phaliales Z.H.Feng, nothosect. nov. — Parentage: Anacamptis sect. Phalaenanthus × Orchis sect. Provinciales.
× Anacamptorchis nothosect. Robustoccatae Z.H.Feng, nothosect. nov. — Parentage: Anacamptis sect. Saccatae × Orchis sect. Robustocalcare.
× Anacamptorchis nothosect. Saccusillae Z.H.Feng, nothosect. nov. — Parentage: Anacamptis sect. Saccatae × Orchis sect. Pusillae.
Cattleya Lindl., Coll. Bot. (Lindley) 7: t. 33 (1824).
Cattleya nothosubgen. Cattmediae Z.H.Feng, nothosubgen. nov. — Parentage: Cattleya subgen. Cattleya × Cattleya subgen. Intermediae.
Cattleya nothosect. Crisleya Z.H.Feng, nothosect. nov. — Parentage: Cattleya sect. Cattleya × Cattleya sect. Crispae.
Cattleya nothoser. Hadroleyleyodes Z.H.Feng, nothoser. nov. — Parentage: Cattleya sect. Cattleya × Cattleya ser. Cattleyodes × Cattleya ser. Hadrolaelia.
Cattleya nothoser. Hadronitis Z.H.Feng, nothoser. nov. — Parentage: Cattleya ser. Hadrolaelia × Cattleya ser. Sophronitis.
Cattleya nothoser. Inteparviodes Z.H.Feng, nothoser. nov. — Parentage: Cattleya subgen. Intermediae × Cattleya ser. Cattleyodes × Cattleya ser. Parviflorae.
Cattleya nothoser. Intesophronitis Z.H.Feng, nothoser. nov. — Parentage: Cattleya subgen. Intermediae × Cattleya ser. Sophronitis.
Cattleya nothoser. Lawolaelia Z.H.Feng, nothoser. nov. — Parentage: Cattleya sect. Lawrenceanae × Cattleya ser. Hadrolaelia.
Cattleya nothoser. Parviodes Z.H.Feng, nothoser. nov. — Parentage: Cattleya sect. Cattleyodes × Cattleya ser. Parviflorae.
× Dactylocamptis P.F.Hunt & Summerh., Watsonia 6: 132 (1965).
× Dactylocamptis nothosect. Andrafoliae Z.H.Feng, nothosect. nov. — Parentage: Anacamptis sect. Andranthus × Dactylorhiza sect. Angustifoliae.
× Dactylocamptis nothosect. Andrarhiza Z.H.Feng, nothosect. nov. — Parentage: Anacamptis sect. Andranthus × Dactylorhiza sect. Dactylorhiza.
× Dactylocamptis nothosect. Andrulatae Z.H.Feng, nothosect. nov. — Parentage: Anacamptis sect. Andranthus × Dactylorhiza sect. Maculatae.
× Dactylocamptis nothosect. Coriorhiza Z.H.Feng, nothosect. nov. — Parentage: Anacamptis sect. Coriophorae × Dactylorhiza sect. Dactylorhiza.
× Dactylocamptis nothosect. Dactylorianthus Z.H.Feng, nothosect. nov. — Parentage: Anacamptis sect. Morianthus × Dactylorhiza sect. Dactylorhiza.
× Dactylocamptis nothosect. Macamptis Z.H.Feng, nothosect. nov. — Parentage: Anacamptis sect. Anacamptis × Dactylorhiza sect. Maculatae.
× Dactylocamptis nothosect. Morculatae Z.H.Feng, nothosect. nov. — Parentage: Anacamptis sect. Morianthus × Dactylorhiza sect. Maculatae.
× Dactylocamptis nothosect. Morifoliae Z.H.Feng, nothosect. nov. — Parentage: Anacamptis sect. Morianthus × Dactylorhiza sect. Angustifoliae.
× Dactylocamptis nothosect. Morucinae Z.H.Feng, nothosect. nov. — Parentage: Anacamptis sect. Morianthus × Dactylorhiza sect. Sambucinae.
× Dactylodenia Garay & H.R.Sweet, Bot. Mus. Leafl. 21: 157 (1966).
× Dactylodenia nothosect. Angustiana Z.H.Feng, nothosect. nov. — Parentage: Dactylorhiza sect. Angustifoliae × Gymnadenia sect. Praegnatiana.
× Dactylodenia nothosect. Balcafoliae Z.H.Feng, nothosect. nov. — Parentage: Dactylorhiza sect. Angustifoliae × Gymnadenia sect. Balcanica.
× Dactylodenia nothosect. Gymnafoliae Z.H.Feng, nothosect. nov. — Parentage: Dactylorhiza sect. Angustifoliae × Gymnadenia sect. Gymnadenia.
× Dactylodenia nothosect. Gymnaglossum (Rolfe) Z.H.Feng, comb. & stat. nov. — Basionym: × Gymnaglossum Rolfe, Orchid Rev. 27: 171 (1919). — Parentage: Dactylorhiza sect. Coeloglossum × Gymnadenia sect. Gymnadenia.
× Dactylodenia nothosect. Mucudenia Z.H.Feng, nothosect. nov. — Parentage: Dactylorhiza sect. Maculatae × Gymnadenia sect. Gymnadenia.
× Dactylodenia nothosect. Mucutiana Z.H.Feng, nothosect. nov. — Parentage: Dactylorhiza sect. Maculatae × Gymnadenia sect. Praegnatiana.
× Dactylodenia nothosect. Samnadenia Z.H.Feng, nothosect. nov. — Parentage: Dactylorhiza sect. Sambucinae × Gymnadenia sect. Gymnadenia.
Dactylorhiza Neck. ex Nevski, Trudy Bot. Inst. Akad. Nauk S.S.S.R., Ser. 1, Fl. Sist. Vyssh. Rast. 4: 332 (1937).
Dactylorhiza nothosect. Anglossum Z.H.Feng, nothosect. nov. — Parentage: Dactylorhiza sect. Angustifoliae × Dactylorhiza sect. Coeloglossum.
× Gymnanacamptis Asch. & Graebn., Syn. Mitteleur. Fl. 3: 854 (1907).
× Gymnanacamptis nothosect. Andradenia Z.H.Feng, nothosect. nov. — Parentage: Anacamptis sect. Andranthus × Gymnadenia sect. Gymnadenia.
× Gymnanacamptis nothosect. Gymorianthus Z.H.Feng, nothosect. nov. — Parentage: Anacamptis sect. Morianthus × Gymnadenia sect. Gymnadenia.
× Orchidactylorhiza Soó & Borsos, Ann. Univ. Sci. Budapest. Rolando Eötvös, Sect. Biol. 8: 315, 317 (1966).
× Orchidactylorhiza nothosect. Anthrofoliae Z.H.Feng, nothosect. nov. — Parentage: Dactylorhiza sect. Angustifoliae × Orchis sect. Anthropophorae.
× Orchidactylorhiza nothosect. Orchifoliae Z.H.Feng, nothosect. nov. — Parentage: Dactylorhiza sect. Angustifoliae × Orchis sect. Orchis.
× Orchidactylorhiza nothosect. Samasculae Z.H.Feng, nothosect. nov. — Parentage: Dactylorhiza sect. Sambucinae × Orchis sect. Masculae.
× Orchigymnadenia E.G.Camus, J. Bot. (Morot) 6: 479 (1892).
× Orchigymnadenia nothosubgen. Madenia Z.H.Feng, nothosubgen. nov. — Parentage: Gymnadenia subgen. Gymnadenia × Orchis subgen. Masculae.
× Orchigymnadenia nothosect. Madenia Z.H.Feng, nothosect. nov. — Parentage: Gymnadenia sect. Gymnadenia × Orchis sect. Masculae.
× Orchimantoglossum Asch. & Graebn., Syn. Mitteleur. Fl. 3: 799 (1907)
× Orchimantoglossum nothosubgen. Barlorchis (Kalop. & Constantin.) Z.H.Feng, comb. & stat. nov. — Basionym: × Barlorchis Kalop. & Constantin., Mitteilungsbl. Arbeitskreis Heimische Orchid. Baden-Württemberg 25(4): 492 (1993). — Parentage: Himantoglossum subgen. Barlia × Orchis subgen. Orchis.
× Orchimantoglossum nothosect. Barlitalicae Z.H.Feng, nothosect. nov. — Parentage: Himantoglossum subgen. Barlia × Orchis sect. Italicae.
× Orchimantoglossum nothosect. Barropophorae Z.H.Feng, nothosect. nov. — Parentage: Himantoglossum subgen. Barlia × Orchis sect. Anthropophorae.
Paphiopedilum Pfitzer, Morph. Stud. Orchideenbl. 11 (1886).
Paphiopedilum nothosubgen. Brachypaphiopedilum Z.H.Feng, nothosubgen. nov. — Parentage: Paphiopedilum subgen. Brachypetalum × Paphiopedilum subgen. Paphiopedilum.
Paphiopedilum nothosect. Brachybarbata Z.H.Feng, nothosect. nov. — Parentage: Paphiopedilum subgen. Brachypetalum × Paphiopedilum sect. Barbata.
Paphiopedilum nothosect. Brachycoryopedilum Z.H.Feng, nothosect. nov. — Parentage: Paphiopedilum subgen. Brachypetalum × Paphiopedilum sect. Coryopedilum.
Paphiopedilum nothosect. Brachypaphiobata Z.H.Feng, nothosect. nov. — Parentage: Paphiopedilum subgen. Brachypetalum × Paphiopedilum sect. Barbata × Paphiopedilum sect. Paphiopedilum.
Paphiopedilum nothosect. Cophiopedilum Z.H.Feng, nothosect. nov. — Parentage: Paphiopedilum sect. Coryopedilum × Paphiopedilum sect. Paphiopedilum.
Phalaenopsis Blume, Bijdr. Fl. Ned. Ind. 7: 294 (1825).
Phalaenopsis nothosect. Polychilopsis Z.H.Feng, nothosect. nov. — Parentage: Phalaenopsis sect. Phalaenopsis × Phalaenopsis sect. Polychilos.
Ranunculaceae
Pulsatilla Mill., Gard. Dict. Abr., ed. 4. [unpaged] (1754).
Pulsatilla nothosubgen. Preotilla Z.H.Feng, nothosubgen. nov. — Parentage: Pulsatilla subgen. Preonanthus (DC.) Zaikov & Shmakov × Pulsatilla subgen. Pulsatilla.
Remarks: The nothosubgeneric name first published by Feng is invalid because one of its parentage is invalidly published, the name is here validated by providing legitimate parentage.
Pulsatilla nothosect. Veranthus Z.H.Feng, nothosect. nov. — Parentage: Pulsatilla sect. Preonanthus × Pulsatilla sect. Vernales.
Pulsatilla nothosect. Versatilla Z.H.Feng, nothosect. nov. — Parentage: Pulsatilla sect. Pulsatilla × Pulsatilla sect. Vernales.
Pulsatilla nothosubsect. Patenses Z.H.Feng, nothosubsect. nov. — Parentage: Pulsatilla sect. Patentes × Pulsatilla subsect. Pratenses.
Pulsatilla nothosubsect. Pratilla Z.H.Feng, nothosubsect. nov. — Parentage: Pulsatilla subsect. Pratenses × Pulsatilla subsect. Pulsatilla.
Pulsatilla nothosubsect. Preotenses Z.H.Feng, nothosubsect. nov. — Parentage: Pulsatilla sect. Preonanthus × Pulsatilla subsect. Pratenses.
Pulsatilla nothosubsect. Preotilla Z.H.Feng, nothosubsect. nov. — Parentage: Pulsatilla sect. Preonanthus × Pulsatilla subsect. Pulsatilla.
Pulsatilla nothosubsect. Vernatilla Z.H.Feng, nothosubsect. nov. — Parentage: Pulsatilla sect. Vernales × Pulsatilla subsect. Pulsatilla.
Pulsatilla nothosubsect. Vertenses Z.H.Feng, nothosubsect. nov. — Parentage: Pulsatilla sect. Vernales × Pulsatilla subsect. Pratenses.
2.3. Nomenclatural Novelties at Specific Level
Aizoaceae
× Delosphyllum schickii (Poind.) Z.H.Feng, comb. nov. — Basionym: Mesembryanthemum × schickii Poind., Cact. Succ. J. (Los Angeles) 6: 51 (1934). — Parentage: Glottiphyllum linguiforme × Delosperma echinatum.
Amaranthaceae
× Osteolaena caput-casuarii (J.H.Willis) Z.H.Feng, comb. nov. — Basionym: Bassia × caput-casuarii J.H.Willis, Vict. Naturalist 73: 153 (1957). — Parentage: Osteocarpum acropterum × Sclerolaena divaricata.
Amaryllidaceae
Narcissus × jalamensis (Patino, Urrutia & J.Valencia) Z.H.Feng, stat. nov. — Basionym: Narcissus × brevitubulosus nothosubsp. jalamensis Patino, Urrutia & J.Valencia, Fl. Montiber. 85: 47 (2023). — Parentage: Narcissus bulbocodium × Narcissus minor.
Narcissus × saenzii (Fern.Casas) Z.H.Feng, stat. nov. — Basionym: Narcissus × alejandrei nothovar. saenzii Fern.Casas, Fontqueria 57(26): 119 (2021). — Parentage: Narcissus gigas × Narcissus jacetanus.
Urceolina × stevensii (N.E.Br.) Z.H.Feng, comb. nov. — Basionym: Eucharis × stevensii N.E.Br., Gard. Chron., ser. 3, 7: 640 (1890). — Parentage: Urceolina candida × Urceolina sanderi.
Apocynaceae
Araujia × intermedia (T.Mey.) Z.H.Feng, comb. nov. — Basionym: Morrenia × intermedia T.Mey., Lilloa 23: 51 (1950). — Parentage: Araujia brachystephana × Araujia odorata.
× Brachypelia nolae-carriae (P.V.Heath) Z.H.Feng, comb. nov. — Basionym: × Brachystemon nolae-carriae P.V.Heath, Calyx 1(1): 20 (1992). — Parentage: Brachystelma barberae × Stapelia similis.
× Huernivalia anemoniflora (Deflers) Z.H.Feng, comb. nov. — Basionym: Stapelia × anemoniflora Deflers, Bull. Soc. Bot. France 43: 118 (1896). — Parentage: Duvalia sulata × Huernia macrocarpa.
Plumeria × angustissima (Urb.) Z.H.Feng, stat. nov. — Basionym: Plumeria × stenopetala nothovar. angustissima Urb., Ark. Bot. 17(7): 50 (1921). — Parentage: Plumeria sericifolia × Plumeria subsessilis.
Stapelia × prostratiflora (L.C.Leach) Z.H.Feng, stat. nov. — Basionym: Stapelia erectiflora var. prostratiflora L.C.Leach, S. African J. Bot. 3: 178 (1984). — Parentage: Stapelia erectiflora × Stapelia glanduliflora.
Asteraceae
× Chrysania konoana (Makino) Z.H.Feng, comb. nov. — Basionym: Chrysanthemum × konoanum Makino, J. Jap. Bot. 2: 18 (1921). — Parentage: Ajania pallasiana × Chrysanthemum makinoi.
× Chrysania marginata (Miq.) Z.H.Feng, comb. nov. — Basionym: Pyrethrum marginatum Miq., Fl. Ned. Ind. 2(1): 85 (1856). — Parentage: Ajania pacifica × [Chrysanthemum × morifolium].
× Chrysania todaiensis (H.Ohashi & Yonek.) Z.H.Feng, comb. nov. — Basionym: Chrysanthemum × todaiense H.Ohashi & Yonek., J. Jap. Bot. 79: 192 (2004). — Parentage: Ajania rupestris × Chrysanthemum makinoi.
× Cicuca favratii (Wilczek) Z.H.Feng, comb. nov. — Basionym: Cicerbita favratii Wilczek, Bull. Soc. Vaud. 51: 333 (1917). — Parentage: Cicerbita alpina × Lactuca plumieri.
× Ozodes purdiei (Petrie) Z.H.Feng, comb. nov. — Basionym: Helichrysum purdiei Petrie, Trans. & Proc. New Zealand Inst. 22: 440 (1890). — Parentage: Anaphalioides bellidioides × Ozothamnus glomeratus.
Pericallis × andersonii (Pépin) Z.H.Feng, comb. nov. — Basionym: Cineraria × andersonii Pépin, Ann. Fl. Pomone 5: 284 (1837). — Parentage: Pericallis aurita × Pericallis cruenta.
Pilosella × astrotricha (Nägeli & Peter) Z.H.Feng, comb. & stat. nov. — Basionym: Hieracium pilosella subsp. astrotrichum Nägeli & Peter, Hierac. Mitt.-Eur. 1: 171 (1885). — Parentage: Pilosella schultesii × Pilosella velutina.
× Psepaurea hybridus (Heydt) Z.H.Feng, comb. nov. — Basionym: × Centaureopappus hybridus Heydt, Möller's Deutsche Gärtn.-Zeitung 47: 166 (1932). — Parentage: Centaurea orientalis × Psephellus pulcherrimus.
Ptilostemon × sprengeri (Wittm.) Z.H.Feng, comb. nov. — Basionym: Chamaepeuce × sprengeri Wittm., Gart.-Zeitung (Berlin) 1: 439 (1882). — Parentage: Ptilostemon casabonae × Ptilostemon diacantha.
Berberidaceae
Berberis × emeiensis (J.M.H.Shaw) Z.H.Feng, comb. nov. — Basionym: Mahonia × emeiensis J.M.H.Shaw, Plantsman, n.s., 10: 46 (2011). — Parentage: Berberis gracilipes × Berberis schochii.
Berberis × lindsayi (Yeo) Z.H.Feng, comb. nov. — Basionym: Mahonia × lindsayi Yeo, J. Roy. Hort. Soc. 97: 38 (1972). — Parentage: Berberis japonica × Berberis napaulensis.
Berberis × savilliana (J.M.H.Shaw) Z.H.Feng, comb. nov. — Basionym: Mahonia × savilliana J.M.H.Shaw, Plantsman, n.s., 10: 46 (2011). — Parentage: Berberis eurybracteata × Berberis gracilipes.
Berberis × wagneri (Jouin) Z.H.Feng, comb. & stat. nov. — Basionym: Mahonia pinnata var. wagneri Jouin, Mitt. Deutsch. Dendrol. Ges. 1910: 90 (1910). — Parentage: Berberis aquifolia × Berberis pinnata.
Boraginaceae
Moltkia × froebelii (Sünd.) Z.H.Feng, comb. nov. — Basionym: Lithospermum × froebelii Sünd., Allg. Bot. Z. Syst. 12: 92 (1906). — Parentage: Moltkia petraea × Moltkia suffruticosa.
Cactaceae
Turbinicarpus × amoenulus nothosubsp. calvescens (Halda & Panar.) Z.H.Feng, comb. & stat. nov. — Basionym: Turbinicarpus × calvescens Halda & Panar., Acta Mus. Richnov., Sect. Nat. 10(2): 167 (2003). — Parentage: Turbinicarpus pseudopectinatus × Turbinicarpus schmiedickeanus subsp. jauernigii.
Turbinicarpus schmiedickeanus nothosubsp. anosmus (Halda & Panar.) Z.H.Feng, comb. & stat. nov. — Basionym: Turbinicarpus × anosmus Halda & Panar., Acta Mus. Richnov., Sect. Nat. 10(2): 163 (2003). — Parentage: Turbinicarpus schmiedickeanus subsp. andersonii × Turbinicarpus schmiedickeanus subsp. bonatzii.
× Turdenicarpus dubeniorum (Halda, Kracík & Malina) Z.H.Feng, comb. nov. — Basionym: Turbinicarpus × dubeniorum Halda, Kracík & Malina, Acta Mus. Richnov., Sect. Nat. 9(1): 75 (2002). — Parentage: Kadenicarpus pseudomacrochele × Turbinicarpus alonsoi.
Crassulaceae
Aeonium × extinctum (P.V.Heath) Z.H.Feng, comb. nov. — Basionym: × Greenonium extinctum P.V.Heath, Calyx 2(2): 59 (1992). — Parentage: Aeonium aureum × Aeonium glutinosum.
Aeonium × lajense (O.Arango) Z.H.Feng, comb. nov. — Basionym: × Greenonium lajense O.Arango, Collect. Bot. (Barcelona) 42(e008): 12 (2023). — Parentage: Aeonium aureum × Aeonium saundersii.
Cupressaceae
Hesperocyparis × symbolica (Maerki) Z.H.Feng, comb. nov. — Basionym: Cupressus × symbolica Maerki, Bull. Cupressus Conservation Proj. 4(2): 79 (2015). — Parentage: Hesperocyparis glabra × Hesperocyparis lusitanica.
Cyperaceae
Carex × flatbergii Z.H.Feng, nom. nov. — Replaced synonym: Carex × lidii Flatberg, Norweg. J. Bot. 19(2): 93 (1972), non Carex × lidii Hadač, Stud. Bot. Čech. 5: 2 (1942). — Parentage: Carex canescens × Carex chordorrhiza.
Carex × tablelandica Z.H.Feng, nom. nov. — Replaced synonym: Carex × elatior (Kük.) K.A.Ford, New Zealand J. Bot. 63(5): 2768 (2025), non Carex elatior Boeckeler, Abh. Naturwiss. Vereins Bremen 7: 41 (1880). — Basionym: Carex kirkii var. elatior Kük., Man. New Zealand Fl. 811 (1906). — Parentage: Carex ambita × Carex trachycarpa.
Equisetaceae
Equisetum × nipponicum (Lubienski) Z.H.Feng, stat. nov. — Basionym: Equisetum moorei nothosubsp. nipponicum Lubienski, Fern Gaz. 21(8): 474 (2022). — Parentage: Equisetum praealtum × Equisetum ramosissimum.
Ericaceae
Arctous × victorinii (Roll.-Germ.) Z.H.Feng, comb. nov. — Basionym: Arctostaphylos × victorinii Roll.-Germ., Fl. Anticosti-Minganie: 463 (1969). — Parentage: Arctous alpinus × Arctous ruber.
Gaultheria × rupicoloides (C.K.Schneid.) Z.H.Feng, comb. nov. — Basionym: Pernettya rupicoloides C.K.Schneid., Ill. Handb. Laubholzk. [C.K.Schneider] 2: 540 (1911). — Parentage: Gaultheria mucronata × Gaultheria myrsinoides.
Fabaceae
× Astoxytropis arnaudii (Petitm.) Z.H.Feng & K.W.Jiang, comb. nov. — Basionym: Oxytropis × arnaudii Petitm., Monde Pl. Rev. Mens. Bot. 7: 16 (1897). — Parentage: Astragalus alpinus × Oxytropis helvetica.
Chamaecytisus × semerenkoanus (Sennikov & Val.N.Tikhom.) Z.H.Feng & K.W.Jiang, comb. nov. — Basionym: Cytisus × semerenkoanus Sennikov & Val.N.Tikhom., PhytoKeys 238: 173 (2024). — Parentage: Chamaecytisus elongatus × Chamaecytisus ruthenicus.
Chamaecytisus × vadasii (J.Wagner) Z.H.Feng & K.W.Jiang, comb. nov. — Basionym: Cytisus × vadasii J.Wagner, Magyar Bot. Lapok 14: 78 (1915). — Parentage: Chamaecytisus austriacus × Chamaecytisus ratisbonensis.
Lotus × bicknellianus (A.Berger & Dinter) Z.H.Feng & K.W.Jiang, comb. nov. — Basionym: Dorycnium × bicknellianum A.Berger & Dinter, Bull. Soc. Bot. Ital. 1910: 137 (1910). — Parentage: Lotus dorycnium × Lotus hirsutus.
Medicago × elmaluensis (Širj.) Z.H.Feng & K.W.Jiang, comb. nov. — Basionym: Trigonella × elmaluensis Širj., Spisy Přír. Fak. Masarykovy Univ. 136: 31 (1931). — Parentage: Medicago crassipes × Medicago monspeliaca.
Gesneriaceae
× Eucongia multiflora (Veitch ex J.Dix) Z.H.Feng, comb. nov. — Basionym: Stenogastra × multiflora Veitch ex J.Dix, Proc. Roy. Hort. Soc. London 2: 373 (1862). — Parentage: Eucodonia verticillata × Sinningia concinna.
Ramonda × jurasekii (Halda) Z.H.Feng, comb. nov. — Basionym: × Jankaemonda jurasekii Halda, Acta Mus. Richnov., Sect. Nat. 19(3-4): 50 (2012). — Parentage: Ramonda heldreichii × Ramonda serbica.
Ramonda × stibicii (Halda) Z.H.Feng, comb. nov. — Basionym: × Jankaemonda stibicii Halda, Acta Mus. Richnov., Sect. Nat. 19(3-4): 50 (2012). — Parentage: Ramonda heldreichii × Ramonda nathaliae.
Ramonda × vandedemii (Halda) Z.H.Feng, comb. nov. — Basionym: × Jankaemonda vandedemii Halda, Preslia 51(4): 375 (1979). — Parentage: Ramonda heldreichii × Ramonda myconi.
Iridaceae
Iris × catharinae (D.Dubovik) Z.H.Feng, comb. nov. — Basionym: Iridodictyum × catharinae D.Dubovik, Fl. Belarusi 3: 209 (2017). — Parentage: Iris histrioides × Iris winogradowii.
Iris × georgiae (D.Dubovik) Z.H.Feng, comb. nov. — Basionym: Iridodictyum × georgiae D.Dubovik, Fl. Belarusi 3: 210 (2017). — Parentage: Iris histrioides × Iris reticulata.
Iris × grodnensis (D.Dubovik) Z.H.Feng, comb. nov. — Basionym: Juno × grodnensis D.Dubovik, Fl. Belarusi 3: 228 (2017). — Parentage: Iris bucharica × Iris graeberiana.
Lamiaceae
Micromeria × tenerifolia (R.H.Willemse) Z.H.Feng, comb. nov. — Basionym: Satureja × tenerifolia R.H.Willemse, Willdenowia 21(1-2): 81 (1991). — Parentage: Micromeria hyssopifolia × Micromeria teneriffae.
Micromeria × tenevaria (R.H.Willemse) Z.H.Feng, comb. nov. — Basionym: Satureja × tenevaria R.H.Willemse, Willdenowia 21(1-2): 81 (1991). — Parentage: Micromeria teneriffae × Micromeria varia.
Marantaceae
Calathea × helenae (Carrière) Z.H.Feng, comb. nov. — Basionym: Maranta × helenae Carrière, Rev. Hort. (Paris) 51: 20 (1879). — Parentage: Calathea eximia × Calathea picturata.
Melanthiaceae
× Pseudillium crockerianum (Halda) Z.H.Feng, comb. nov. — Basionym: Trillium × crockerianum Halda, Acta Mus. Richnov., Sect. Nat. 6(3): 234 (1999). — Parentage: Pseudotrillium rivale × Trillium ovatum. — Heterotypic synonym: Trillium × klineanum Halda, Acta Mus. Richnov., Sect. Nat. 6(3): 234 (1999), syn. nov.
Orchidaceae
× Anacamptorchis bonnieriana (E.G.Camus) Z.H.Feng & J.M.H.Shaw, comb. nov. — Basionym: Orchis × bonnieriana E.G.Camus, J. Bot. (Morot) 6: 416 (1892). — Parentage: Anacamptis palustris × Orchis militaris.
× Batepalum ballii (Rolfe) Z.H.Feng & J.M.H.Shaw, comb. nov. — Basionym: Zygopetalum ballii Rolfe, Gard. Chron. ser. 3, 27: 149 (1900). — Parentage: Batemannia colleyi × Zygosepalum lindeniae.
Cattleya × andreana (Maron) Z.H.Feng, comb. nov. — Basionym: × Laeliocattleya andreana Maron, Rev. Hort. (Paris) 67: 401 (1895). — Parentage: Cattleya bicolor × [Cattleya × elegans].
Cattleya × aurora (Rolfe) Z.H.Feng, comb. nov. — Basionym: × Laeliocattleya aurora Rolfe, Gard. Chron. ser. 3, 6: 380 (1889). — Parentage: Cattleya bicalhoi × Cattleya loddigesii.
Cattleya × batemanniana (Rchb.f.) Z.H.Feng, comb. nov. — Basionym: Laelia × batemanniana Rchb.f., Gard. Chron. n.s., 26: 263 (1886). — Parentage: Cattleya coccinea × Cattleya intermedia.
Cattleya × bella (Rchb.f.) Z.H.Feng, comb. nov. — Basionym: Laelia × bella Rchb.f., Gard. Chron. n.s., 21: 174 (1884). — Parentage: Cattleya labiata × Cattleya purpurata. — Heterotypic synonym: × Laeliocattleya arnoldiana Rolfe, J. Orchidées 2: 134 (1891), syn. nov.
Cattleya × broomfieldiensis (Mast.) Z.H.Feng, comb. nov. — Basionym: × Laeliocattleya broomfieldiensis Mast., J. Orchidées 2: 134 (1891). — Parentage: Cattleya dowiana × Cattleya praestans.
Cattleya × callistoglossa (Rchb.f.) Z.H.Feng, comb. nov. — Basionym: Laelia × callistoglossa Rchb.f., Gard. Chron. n.s., 17: 76 (1882). — Parentage: Cattleya coccinea × Cattleya harrisoniana. — Heterotypic synonym: × Sophrocattleya calypso Rolfe, Gard. Chron., ser. 3, 8: 588 (1890), syn. nov.
Cattleya × caloglossa (Rchb.f.) Z.H.Feng, comb. nov. — Basionym: Laelia × caloglossa Rchb.f., Gard. Chron. n.s., 7: 202 (1877). — Parentage: Cattleya labiata × Cattleya lobata.
Cattleya × canhamiana (Rchb.f.) Z.H.Feng, comb. nov. — Basionym: Laelia × canhamiana Rchb.f., Gard. Chron. n.s., 24: 6 (1885). — Parentage: Cattleya mossiae × Cattleya purpurata.
Cattleya × cappei (Anon.) Z.H.Feng, comb. nov. — Basionym: × Laeliocattleya cappei Anon., Orchid Rev. 7: 46 (1899). — Parentage: Cattleya cinnabarina × Cattleya warscewiczii.
Cattleya × cassiope (Rolfe) Z.H.Feng, comb. nov. — Basionym: × Laeliocattleya cassiope Rolfe, Gard. Chron., ser. 3, 6: 620 (1889). — Parentage: [Cattleya × exoniensis] × Cattleya pumila.
Cattleya × decia (Mast.) Z.H.Feng, comb. nov. — Basionym: × Laeliocattleya decia Mast., Gard. Chron., ser. 3, 16: 699 (1894). — Parentage: Cattleya dowiana × Cattleya perrinii.
Cattleya × euterpe (Rchb.f.) Z.H.Feng, comb. nov. — Basionym: Laelia × euterpe Rchb.f., Gard. Chron. ser. 3, 4: 533 (1888). — Parentage: Cattleya bicalhoi × Cattleya crispa.
Cattleya × haroldiana (Jekyll & E.T.Cook) Z.H.Feng, comb. nov. — Basionym: × Laeliocattleya haroldiana Jekyll & E.T.Cook, Garden (London, 1871-1927) 60: 158 (1901). — Parentage: Cattleya dowiana × Cattleya tenebrosa × Cattleya warscewiczii.
Cattleya × jacobiana (Truff.) Z.H.Feng, comb. nov. — Basionym: × Laeliocattleya jacobiana Truff., Rev. Hort. (Paris) 67: 359 (1895). — Parentage: Cattleya mendelii × Cattleya purpurata.
Cattleya × juvenilis (Bleu) Z.H.Feng, comb. nov. — Basionym: Laelia × juvenilis Bleu, J. Orchidées 1: 230 (1890). — Parentage: Cattleya perrinii × Cattleya pumila.
Cattleya × kadina (W.H.Cox) Z.H.Feng, comb. nov. — Basionym: × Laeliocattleya kadina W.H.Cox, Orchid Rev. 29: 98 (1921). — Parentage: [Cattleya × elegans] × Cattleya labiata.
Cattleya × laeta (Mast.) Z.H.Feng, comb. nov. — Basionym: × Sophrocattleya laeta Mart., Gard. Chron. ser. 3, 16: 476 (1894). — Parentage: Cattleya bicalhoi × Cattleya coccinea.
Cattleya × maynardii (Sander ex Carrière & André) Z.H.Feng, comb. nov. — Basionym: × Laeliocattleya maynardii Sander ex Carrière & André, Rev. Hort. (Paris) 65: 294 (1893). — Parentage: Cattleya bicalhoi × Cattleya loddigesii × Cattleya walkeriana.
Cattleya × mylamiana (Rchb.f.) Z.H.Feng, comb. nov. — Basionym: Laelia × mylamiana Rchb.f., Gard. Chron. n.s., 6: 740 (1876). — Parentage: Cattleya crispa × Cattleya granulosa.
Cattleya × ochracea (Grignan) Z.H.Feng, comb. nov. — Basionym: × Laeliocattleya ochracea Grignan, Jardin 15: 7 (1901). — Parentage: Cattleya harpophylla × [Cattleya × sallieri].
Cattleya × oweniae (L.Linden) Z.H.Feng, comb. nov. — Basionym: Laelia × oweniae L.Linden, Lindenia 8: 79 (1892). — Parentage: [Cattleya × elegans] × Cattleya perrinii.
Cattleya × philbrickiana (Rchb.f.) Z.H.Feng, comb. nov. — Basionym: Laelia × philbrickiana Rchb.f., Gard. Chron. n.s., 12: 102 (1879). — Parentage: Cattleya aclandiae × Cattleya purpurata × Cattleya tigrina.
Cattleya × phoebe (O'Brien) Z.H.Feng, comb. nov. — Basionym: × Laeliocattleya phoebe O'Brien, Gard. Chron., ser. 3, 11: 791 (1892). — Parentage: Cattleya mossiae × Cattleya purpurata.
Cattleya × proserpini (Rolfe ex B.S.Williams) Z.H.Feng, comb. nov. — Basionym: × Laeliocattleya proserpini Rolfe ex B.S.Williams, Orch.-Grow. Man., ed. 7: 461 (1893). — Parentage: Cattleya bicalhoi × Cattleya velutina.
Cattleya × sallieri (Maron) Z.H.Feng, comb. nov. — Basionym: × Laeliocattleya sallieri Maron, Rev. Hort. (Paris) 67: 511 (1895). — Parentage: Cattleya loddigesii × Cattleya purpurata.
Cattleya × sedenii (Maron) Z.H.Feng, comb. nov. — Basionym: Laelia × sedenii Rchb.f., Gard. Chron. n.s., 8: 424 (1877). — Parentage: [Cattleya × elegans] × Cattleya violacea.
Cattleya × stella (Rolfe) Z.H.Feng, comb. nov. — Basionym: × Laeliocattleya stella Rolfe, Gard. Chron., ser. 3, 6: 322 (1889). — Parentage: Cattleya crispa × Cattleya purpurata × Cattleya tigrina.
Cattleya × timora (N.E. Br.) Z.H.Feng, comb. nov. — Basionym: Laelia × timora N.E.Br., Gard. Chron., ser. 3, 2: 428 (1887). — Parentage: Cattleya bicalhoi × Cattleya lueddemanniana.
Cattleya × vitellina (O'Brien) Z.H.Feng, comb. nov. — Basionym: Laelia × vitellina O'Brien, Gard. Chron., ser. 3, 13: 365 (1893) — Parentage: Cattleya harpophylla × Cattleya perrinii.
Cymbidium × fowlerae (Schltr.) Z.H.Feng, comb. nov. — Basionym: Cyperorchis × fowlerae Schltr., Repert. Spec. Nov. Regni Veg. 20: 108 (1924). — Parentage: Cymbidium elegans × Cymbidium iridioides.
× Dactylodenia alto-ovirensis (Jakely & Könighofer) Z.H.Feng & J.M.H.Shaw, comb. nov. — Basionym: × Dactylitella alto-ovirensis Jakely & Könighofer, Ber. Arbeitskreis. Heimische Orchid. 35(1): 216 (2018). — Parentage: Dactylorhiza fuchsii × Gymnadenia lithopolitanica.
Dactylorhiza × mithridatea J.M.H.Shaw & Z.H.Feng, nom. nov. — Replaced synonym: × Dactyloglossum ponticum Kohlmüller, Ber. Arbeitskreis. Heimische Orchid. 16(1): 74 (1999), non Dactylorhiza pontica (H.Fleischm. & Hand.-Mazz.) P.Delforge, Naturalistes Belges 81(4): 397 (2000). — Parentage: Dactylorhiza urvilleana × Dactylorhiza viridis.
× Dossisia dominyi (H.J.Veitch) Z.H.Feng & J.M.H.Shaw, comb. nov. — Basionym: Goodyera × dominyi H.J.Veitch, Gard. Chron. 1861: 531 (1861). — Homotypic synonym: Dossinimaria × dominyi (H.J.Veitch) Rolfe, J. Linn. Soc., Bot. 24: 170 (1888). — Parentage: Dossinia marmorata × Ludisia discolor.
× Epilaelia hardyana (Mast.) Z.H.Feng, comb. nov. — Basionym: × Laeliocattleya hardyana Mast., Gard. Chron., ser. 3, 17: 248 (1895). — Parentage: Epidendrum ciliare × Laelia anceps.
× Galeosepalum marginatum (Garay) Z.H.Feng & J.M.H.Shaw, comb. nov. — Basionym: Zygosepalum marginatum Garay, Revista Soc. Colomb. Orquideol. 2(3): 8 (1967). — Parentage: Galeottia grandiflora × Zygosepalum lindeniae.
× Ludochilus dominyi (H.J.Veitch) Z.H.Feng & J.M.H.Shaw, comb. nov. — Basionym: Anoectochilus × dominyi H.J.Veitch, Proc. Roy. Hort. Soc. London 5: 139 (1865). — Homotypic synonym: × Anoectomaria dominyi (H.J.Veitch) Rolfe, J. Linn. Soc., Bot. 24: 170 (1888). — Parentage: Anoectochilus roxburghii × Ludisia discolor.
Paphiopedilum × buchananianum (Rolfe) Z.H.Feng, comb. nov. — Basionym: Cypripedium × buchananianum Rolfe, Gard. Chron. ser. 3, 7: 73 (1890). — Parentage: Paphiopedilum druryi × Paphiopedilum spicerianum.
Paphiopedilum × debosscherianum (Desbois) Z.H.Feng, comb. nov. — Basionym: Cypripedium × debosscherianum Desbois, Rev. Hort. Belge Étrangère 18: 6 (1892). — Parentage: Paphiopedilum spicerianum × Paphiopedilum superbiens.
Paphiopedilum × denisianum (O'Brien) Z.H.Feng, comb. nov. — Basionym: Cypripedium × denisianum O'Brien, Gard. Chron. ser. 3, 16: 118 (1894). — Parentage: [Paphiopedilum × selligerum] × Paphiopedilum superbiens.
Paphiopedilum × fowlerianum (Sander) Z.H.Feng, comb. nov. — Basionym: Cypripedium × fowlerianum Sander, Gard. Chron. ser. 3, 17: 337 (1895). — Parentage: Paphiopedilum bellatulum × [Paphiopedilum × harrisianum].
Paphiopedilum × greyanum (Pitcher & Manda) Z.H.Feng, comb. nov. — Basionym: Cypripedium × greyanum Pitcher & Manda, Proc. Annual Conv. Soc. Amer. Florists 8: 81 (1892). — Parentage: Paphiopedilum ciliolare × Paphiopedilum druryi.
Paphiopedilum × harrisianum (Rchb.f.) Z.H.Feng, comb. nov. — Basionym: Cypripedium × harrisianum Rchb.f., Gard. Chron. 1869: 108 (1869). — Parentage: Paphiopedilum barbatum × Paphiopedilum villosum.
Paphiopedilum × hephaestus (A.H.Kent) Z.H.Feng, comb. nov. — Basionym: Cypripedium × hephaestus A.H.Kent, Gard. Chron. ser. 3, 3: 425 (1888). — Parentage: Paphiopedilum barbatum × Paphiopedilum lawrenceanum.
Paphiopedilum × inspirator (Kerch.) Z.H.Feng, comb. nov. — Basionym: Cypripedium × inspirator Kerch., Rev. Hort. Belge Étrangère 20: 25 (1894). — Parentage: Paphiopedilum spicerianum × Paphiopedilum tonsum.
Paphiopedilum × leda (Mast.) Z.H.Feng, comb. nov. — Basionym: Cypripedium × leda Mast., Gard. Chron. ser. 3, 11: 202 (1892). — Parentage: [Paphiopedilum × harrisianum] × Paphiopedilum venustum.
Paphiopedilum × leysenianum (Desbois) Z.H.Feng, comb. nov. — Basionym: Cypripedium × leysenianum Desbois, Rev. Hort. Belge Étrangère 20: 169 (1894). — Parentage: Paphiopedilum barbatum × Paphiopedilum bellatulum.
Paphiopedilum × luridum (Pitcher & Manda) Z.H.Feng, comb. nov. — Basionym: Cypripedium × luridum Pitcher & Manda, Proc. Annual Conv. Soc. Amer. Florists 8: 81 (1892). — Parentage: Paphiopedilum lawrenceanum × Paphiopedilum villosum.
Paphiopedilum × meirax (Rchb.f.) Z.H.Feng, comb. nov. — Basionym: Cypripedium × meirax Rchb.f., Gard. Chron. n.s. 14: 524 (1880). — Parentage: Paphiopedilum fairrieanum × Paphiopedilum venustum.
Paphiopedilum × pallas (N.E.Br.) Z.H.Feng, comb. nov. — Basionym: Cypripedium × pallas N.E.Br., Gard. Chron. ser. 3, 9: 360 (1891). — Parentage: Paphiopedilum callosum × [Paphiopedilum × calophyllum].
Paphiopedilum × pallens (Pitcher & Manda) Z.H.Feng, comb. nov. — Basionym: Cypripedium × pallens Pitcher & Manda, Proc. Annual Conv. Soc. Amer. Florists 8: 81 (1892). — Parentage: Paphiopedilum dayanum × Paphiopedilum spicerianum.
Paphiopedilum × patens (Rchb.f.) Z.H.Feng, comb. nov. — Basionym: Cypripedium × patens Rchb.f., Gard. Chron. n.s. 8: 456 (1877). — Parentage: Paphiopedilum barbatum × Paphiopedilum hookerae.
Paphiopedilum × pavoninum (Rchb.f.) Z.H.Feng, comb. nov. — Basionym: Cypripedium × pavoninum Rchb.f., Gard. Chron. ser. 3, 3: 264 (1888). — Parentage: Paphiopedilum venustum × Paphiopedilum villosum var. boxallii.
Paphiopedilum × peruzzisnum (Recenti) Z.H.Feng, comb. nov. — Basionym: Cypripedium × peruzzisnum Recenti, Bull. Soc. Tosc. Ortic. ser. 4, 9: 1 (1924). — Parentage: Paphiopedilum haynaldianum × Paphiopedilum spicerianum.
Paphiopedilum × pleistochlorum (Rchb.f.) Z.H.Feng, comb. nov. — Basionym: Cypripedium × pleistochlorum Rchb.f., Gard. Chron. ser. 3, 2: 552 (1887). — Parentage: Paphiopedilum javanicum × Paphiopedilum superbiens.
Paphiopedilum × poyntzianum (O'Brien) Z.H.Feng, comb. nov. — Basionym: Cypripedium × poyntzianum O'Brien, Gard. Chron. ser. 3, 15: 36 (1894). — Parentage: Paphiopedilum callosum × Paphiopedilum hookerae.
Paphiopedilum × rolfei (Jekyll & E.T.Cook) Z.H.Feng, comb. nov. — Basionym: Cypripedium × rolfei Jekyll & E.T.Cook, Garden (London) 60: 158 (1901). — Parentage: Paphiopedilum bellatulum × Paphiopedilum rothschildianum.
Paphiopedilum × selligerum (H.J.Veitch) Z.H.Feng, comb. nov. — Basionym: Cypripedium × selligerum H.J.Veitch, Gard. Chron. n.s. 3: 728 (1875). — Parentage: Paphiopedilum barbatum × Paphiopedilum philippinense.
Paphiopedilum × thayerianum (Sander) Z.H.Feng, comb. nov. — Basionym: Cypripedium × thayerianum Sander, Gard. Chron. ser. 3, 14: 192 (1893). — Parentage: Paphiopedilum lawrenceanum × Paphiopedilum villosum var. boxallii.
Paphiopedilum × triumphans (O'Brien) Z.H.Feng, comb. nov. — Basionym: Cypripedium × triumphans O'Brien, Gard. Chron. ser. 3, 15: 198 (1894). — Parentage: [Paphiopedilum × harrisianum] × [Paphiopedilum × oenanthum].
Paphiopedilum × umlauftianum (Sander) Z.H.Feng, comb. nov. — Basionym: Cypripedium × umlauftianum Sander, Gard. Chron. ser. 3, 14: 70 (1893). — Parentage: Paphiopedilum insigne × Paphiopedilum lawrenceanum.
Paphiopedilum × vernixium (Rchb.f.) Z.H.Feng, comb. nov. — Basionym: Cypripedium × vernixium Rchb.f., Gard. Chron. n.s. 11: 398 (1879). — Parentage: Paphiopedilum argus × Paphiopedilum villosum.
Paphiopedilum × vervaetianum (Rchb.f.) Z.H.Feng, comb. nov. — Basionym: Cypripedium × vervaetianum Rchb.f., Gard. Chron. ser. 3, 3: 712 (1888). — Parentage: Paphiopedilum lawrenceanum × Paphiopedilum superbiens.
Paphiopedilum × wellesleyanum (H.J.Veitch) Z.H.Feng, comb. nov. — Basionym: Cypripedium × wellesleyanum H.J.Veitch, Gard. Chron. ser. 3, 35: 270 (1904). — Parentage: Paphiopedilum bellatulum × Paphiopedilum concolor.
Papilionanthe × joaquimiae (Rolfe & Hurst) Z.H.Feng, comb. nov. — Basionym: Vanda × joaquimiae Rolfe & Hurst, Orchid Stud-Book 236 (1909). — Parentage: Papilionanthe hookeriana × Papilionanthe teres.
Phragmipedium × finetianum (Sander) Z.H.Feng, comb. nov. — Basionym: Selenipedium × finetianum Sander, Orchid Rev. 3: 102 (1895). — Parentage: Phragmipedium caudatum × [Phragmipedium × sedenii].
Phragmipedium × nitidissimum (Rchb.f.) Z.H.Feng, comb. nov. — Basionym: Cypripedium × nitidissimum Rchb.f., Gard. Chron. ser. 3, 4: 6 (1888). — Parentage: Phragmipedium caricinum × Phragmipedium caudatum × Phragmipedium longifolium.
× Serapicamptis garriuscola (Soca) Z.H.Feng, comb. nov. — Basionym: × Herorchiserapias garriuscola Soca, Carnets Bot. 265: 12 (2025). — Parentage: Anacamptis palustris × Serapias vomeracea.
Plantaginaceae
Globularia × gomezcasaresii (Carlón, M.Laínz, Moreno Mor., Rodr.Berd. & Ó.Sánchez) Z.H.Feng, stat. nov. — Basionym: Globularia × montiberica nothosubsp. gomezcasaresii Carlón, M.Laínz, Moreno Mor., Rodr.Berd. & Ó.Sánchez, Doc. Jard. Bot. Atlántico 10: 86 (2014). — Parentage: Globularia repens × Globularia vulgaris.
Poaceae
Puccinellia × burdonii (Druce) Z.H.Feng, comb. nov. — Basionym: Glyceria × burdoni Druce, Rep. Bot. Soc. Exch. Club Brit. Isles 5(1): 61 (1917). — Parentage: Puccinellia fasciculata × Puccinellia maritima.
Sporobolus × onubensis (Sánchez Gullón, M.D.Infante & B.Gallego) Z.H.Feng, comb. nov. — Basionym: Spartina × onubensis Sánchez Gullón, M.D.Infante & B.Gallego, Pl. Syst. Evol. 305: 544 (2019). — Parentage: Sporobolus maritimus × Sporobolus montevidensis.
× Thinoelymus oliveri (Druce) Z.H.Feng, comb. nov. — Basionym: Agropyron × oliveri Druce, Bot. Exch. Club Brit. Isles Rep. 3(1): 38 (1912). — Parentage: Elymus repens × Thinopyrum pycnanthum.
Polypodiaceae
Pleopeltis × haitiensis Z.H.Feng, nom. nov. — Replaced synonym:—Pleopeltis × cuspidata (Desv.) Boudrie, Cremers & Viane, J. Bot. Soc. Bot. France 103(1): 23 (2022), non Pleopeltis cuspidata C.Presl, Tent. Pterid. 193 (1836). — Basionym: Aspidium cuspidatum Desv., Mém. Soc. Linn. Paris 6(3): 244 (1827). — Parentage: Pleopeltis macrocarpa × Pleopeltis thyssanolepis.
Primulaceae
Evotrochis × jenensis (O.Schwarz) Z.H.Feng, comb. nov. — Basionym: Primula × jenensis O.Schwarz, Wiss. Z. Friedrich-Schiller-Univ. Jena, Math.-Naturwiss. Reihe 21(5-6): 967 (1972). — Parentage: Evotrochis edelbergii × Evotrochis verticillata.
Ranunculaceae
× Rankwithia aconitoides (Rouy) Z.H.Feng, comb. nov. — Basionym: Ranunculus × aconitoides Rouy, Ill. Pl. Eur. 4: 25 (1895). — Parentage: Beckwithia glacialis × Ranunculus aconitifolius.
× Rankwithia algidus (Brügger) Z.H.Feng, comb. nov. — Basionym: Ranunculus × algidus Brügger, Jahresber. Naturf. Ges. Graubündens 2(23-24): 78 (1880). — Parentage: Beckwithia glacialis × Ranunculus alpestris.
Ranunculus × udmurtiensis Z.H.Feng, nom. nov. — Replaced synonym: Ranunculus × algidus (Kapit.) A.A.Bobrov, Phytotaxa 319(1): 23 (2017), non Ranunculus × algidus Brügger, Jahresber. Naturf. Ges. Graubündens 2(23-24): 78 (1880). — Basionym: Batrachium algidum Kapit., Bot. Zhurn. (Moscow & Leningrad) 91(2): 277 (2006). — Parentage: Ranunculus kauffmannii × Ranunculus mongolicus.
Rosaceae
Aria × pseudoporrigentiformis (T.C.G.Rich & L.Houston) Z.H.Feng, comb. nov. — Basionym: Sorbus × pseudoporrigentiformis T.C.G.Rich & L.Houston, Brit. Irish Bot. 7(1): 31 (2025). — Parentage: Aria edulis × Aria porrigentiformis.
× Tormaria kitaibeliana (Baksay & Kárpát) Z.H.Feng, comb. nov. — Basionym: Sorbus × kitaibeliana Baksay & Kárpáti, Feddes Repert. Spec. Nov. Regni Veg. 62: 297 (1960). — Parentage: Aria danubialis × Torminalis glaberrima.
Saxifragaceae
× Mitarella grayana (Engl.) Z.H.Feng, comb. nov. — Basionym: Mitella × grayana Engl., Nat. Pflanzenfam., ed. 2 [Engler & Prantl] 18a: 161 (1930). — Parentage: Mitella diphylla × Tiarella cordifolia.
× Mitarella tiarelloides (Engl.) Z.H.Feng, comb. nov. — Basionym: Mitella × tiarelloides Engl., Nat. Pflanzenfam., ed. 2 [Engler & Prantl] 18a: 161 (1930). — Parentage: Mitella nuda × Tiarella cordifolia.
2.4. Nomenclatural Novelties at Infraspecific Level
Amaryllidaceae
Narcissus bulbocodium nothovar. abulensis (Fern.Casas) Z.H.Feng, comb. & stat. nov. — Basionym: Narcissus × abulensis Fern.Casas, Fontqueria 56(52): 554 (2016). — Parentage: Narcissus bulbocodium var. bulbocodium × Narcissus bulbocodium var. minor.
Narcissus × ernii nothovar. guadalupensis (Fern.Casas) Z.H.Feng, comb. & stat. nov. — Basionym: Narcissus × guadalupensis Fern.Casas, Fontqueria 57(26): 109 (2021). — Parentage: Narcissus bulbocodium var. minor × Narcissus triandrus subsp. triandrus.
Narcissus × jalamensis nothosubsp. sanabrensis (Patino, Urrutia & J.Valencia) Z.H.Feng, comb. nov. — Basionym: Narcissus × brevitubulosus nothosubsp. sanabrensis Patino, Urrutia & J.Valencia, Fl. Montiber. 85: 47 (2023). — Parentage: Narcissus bulbocodium subsp. quintanilhae × Narcissus minor.
Apiaceae
Pimpinella saxifraga nothosubsp. subnigra (Tzvelev) Z.H.Feng, comb. & stat. nov. — Basionym: Pimpinella × subnigra Tzvelev, Novosti Sist. Vyssh. Rast. 33: 198 (2001). — Parentage: Pimpinella saxifraga subsp. nigra × Pimpinella saxifraga subsp. saxifraga.
Cistaceae
Fumana ericoides nothosubsp. saguntina (Pérez Dacosta) Z.H.Feng, comb. & stat. nov. — Basionym: Fumana × saguntina Pérez Dacosta, Flora Montiber. 51: 127 (2012). — Parentage: Fumana ericoides subsp. ericoides × Fumana ericoides subsp. montana.
Helianthemum apenninum nothosubsp. serranicum (Mateo) Z.H.Feng, comb. & stat. nov. — Basionym: Helianthemum × serranicum Mateo, Fl. Montiber. 50: 33 (2012). — Parentage: Helianthemum apenninum subsp. apenninum × Helianthemum apenninum subsp. stoechadifolium.
Helianthemum cinereum nothosubsp. pseudocinereum (Pérez Dacosta & Mateo) Z.H.Feng, comb. & stat. nov. — Basionym: Helianthemum × pseudocinereum Pérez Dacosta & Mateo, Fl. Montiber. 50: 50 (2012). — Parentage: Helianthemum cinereum subsp. cinereum × Helianthemum cinereum subsp. rotundifolium.
Crassulaceae
Crassula alpestris nothosubsp. acocksii (P.V.Heath) Z.H.Feng, comb. & stat. nov. — Basionym: Tetraphyle alpestris nothovar. acocksii P.V.Heath, Calyx 3(3): 107 (1993). — Parentage: Crassula alpestris subsp. alpestris × Crassula alpestris subsp. massonii.
Fabaceae
Anthyllis vulneraria nothosubsp. polyphylloides (Juz.) Z.H.Feng & K.W.Jiang, comb. & stat. nov. — Basionym: Anthyllis × polyphylloides Juz., Fl. URSS 11: 276, 393 (1945). — Parentage: Anthyllis vulneraria subsp. lapponica × Anthyllis vulneraria subsp. polyphylla.
Lamiaceae
Clinopodium nepeta nothosubsp. padureanum (Rottenst.) Z.H.Feng, comb. & stat. nov. — Basionym: Clinopodium × padureanum Rottenst., Joannea Bot. 18: 134 (2022). — Parentage: Clinopodium nepeta subsp. spruneri × Clinopodium nepeta subsp. subisodontum.
Mentha spicata nothosubsp. vergoereri (Starm.) Z.H.Feng, comb. & stat. nov. — Basionym: Mentha × vergoereri Starm., Fritschiana 68: 10 (2011). — Parentage: Mentha spicata subsp. condensata × Mentha spicata subsp. spicata.
Salvia officinalis nothosubsp. albaladejitoi (G.Ortiz, Herráiz, García Cardo, Melero & Raúl Sánchez) Z.H.Feng, comb. nov. — Basionym: Salvia × accidentalis nothosubsp. albaladejitoi G.Ortiz, Herráiz, García Cardo, Melero & Raúl Sánchez, Fl. Montiber. 87: 68 (2023). — Parentage: Salvia officinalis subsp. lavandulifolia × Salvia officinalis subsp. officinalis.
Orchidaceae
× Anacamptorchis paschae nothosubsp. cretensis J.M.H.Shaw & Z.H.Feng, nom. nov. — Replaced synonym: Orchis × paschae Hautz., Ann. Naturhist. Mus. Wien 81: 69 (1978). — Parentage: Anacamptis collina × Orchis spitzelii subsp. nitidifolia.
Dactylorhiza × braunii nothosubsp. robustior (Hartm.) Z.H.Feng, comb. & stat. nov. — Basionym: Orchis latifolia var. robustior Hartm., Handb. Skand. Fl. [ed. 3] 204 (1838). — Parentage: Dactylorhiza fuchsii × Dactylorhiza majalis subsp. elatior.
Dactylorhiza × dufftiana nothosubsp. kuuskiae (E.Breiner & R.Breiner) Z.H.Feng, comb. & stat. nov. — Basionym: Dactylorhiza × kuuskiae E.Breiner & R.Breiner, Ber. Arbeitskreis. Heimische Orchid. 15(2): 34 (1998). — Parentage: Dactylorhiza lapponica × Dactylorhiza majalis subsp. baltica.
Epipactis helleborine nothosubsp. soguksuensis (Kreutz) Z.H.Feng, comb. & stat. nov. — Basionym: Epipactis × soguksuensis Kreutz, Eurorchis 9: 7 (1997). — Parentage: Epipactis helleborine subsp. helleborine × Epipactis helleborine subsp. tremolsii.
Epipactis × schmalhausenii nothosubsp. gauckleri (J.Weigelt & Riech.) Z.H.Feng & J.M.H. Shaw, comb. & stat. nov. — Basionym: Epipactis × gauckleri J.Weigelt & Riech., Ber. Arbeitskreis. Heimische Orchid. 19(1): 114, 112 (2002). — Parentage: Epipactis atrorubens × Epipactis helleborine subsp. orbicularis.
Epipactis × stephensonii nothosubsp. lachmanii (Lachmair) Z.H.Feng & J.M.H. Shaw, comb. & stat. nov. — Basionym: Epipactis × lachmanii Lachmair, Stapfia 112: 91 (2021). — Parentage: Epipactis helleborine subsp. orbicularis × Epipactis leptochila subsp. neglecta.
Ophrys × albertiana nothosubsp. angiullii (D'Alonzo) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × angiullii D'Alonzo, GIROS Orch. Spont. Eur. 67(2): 154 (2024). — Parentage: Ophrys apifera × Ophrys holosericea subsp. oxyrrhynchos.
Ophrys × apicula nothosubsp. alejandrei (Benito) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × alejandrei Benito, Estud. Mus. Cienc. Nat. Álava 23: 57 (2011). — Parentage: Ophrys insectifera × Ophrys sphegodes subsp. atrata.
Ophrys × apicula nothosubsp. barbieriarum (Romolini & Soca) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × barbieriarum Romolini & Soca, GIROS Orch. Spont. Eur. 58: 380 (2015). — Parentage: Ophrys insectifera × Ophrys sphegodes subsp. passionis.
Ophrys × apicula nothosubsp. rasinii (Soca) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × rasinii Soca, J. Eur. Orch. 49: 379 (2017). — Parentage: Ophrys insectifera × Ophrys sphegodes subsp. sphegodes.
Ophrys × arachnitiformis nothosubsp. broeckii (A.Camus) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × broeckii A.Camus, Rivièra Sci. 11: 60 (1924). — Parentage: Ophrys holosericea × Ophrys sphegodes subsp. araneola × Ophrys sphegodes subsp. sphegodes.
Ophrys × arachnitiformis nothosubsp. buffaloriae (Romolini & Soca) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × buffaloriae Romolini & Soca, GIROS Notiz. 55: 57 (2014). — Parentage: Ophrys holosericea subsp. parvimaculata × Ophrys sphegodes subsp. atrata.
Ophrys × arachnitiformis nothosubsp. carnazzae (Galesi) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × carnazzae Galesi, J. Eur. Orch. 31: 309 (1999). — Parentage: Ophrys holosericea × Ophrys sphegodes subsp. atrata.
Ophrys × arachnitiformis nothosubsp. centomoi (D'Alonzo & Perilli) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × centomoi D'Alonzo & Perilli, GIROS Orch. Spont. Eur. 64: 68 (2021). — Parentage: Ophrys holosericea subsp. holoserica × Ophrys sphegodes subsp. passionis.
Ophrys × arachnitiformis nothosubsp. coturii (Romolini & Soca) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × coturii Romolini & Soca, GIROS Notiz. 55: 56 (2014). — Parentage: Ophrys holosericea subsp. apulica × Ophrys sphegodes subsp. passionis.
Ophrys × arachnitiformis nothosubsp. coulotii (Soca) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × coulotii Soca, Monde Pl. 91: 15 (1997). — Parentage: Ophrys holosericea subsp. parvimaculata × Ophrys sphegodes subsp. passionis.
Ophrys × arachnitiformis nothosubsp. daissiorum (H.Baumann, Giotta, Künkele, R.Lorenz & Piccitto) Z.H.Feng, comb. nov. — Basionym: Ophrys × maladroxiensis nothosubsp daissiorum H.Baumann, Giotta, Künkele, R.Lorenz & Piccitto, J. Eur. Orch. 27: 223 (1995). — Parentage: Ophrys holosericea subsp. chestermanii × Ophrys holosericea subsp. holosericea × Ophrys sphegodes.
Ophrys × arachnitiformis nothosubsp. difonzoi (D'Alonzo & Perilli) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × difonzoi D'Alonzo & Perilli, GIROS Orch. Spont. Eur. 59: 300 (2016). — Parentage: Ophrys holosericea subsp. parvimaculata × Ophrys sphegodes subsp. sphegodes.
Ophrys × arachnitiformis nothosubsp. franciniae (Bianco, Medagli, D'Emerico & Ruggiero) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × franciniae Bianco, Medagli, D'Emerico & Ruggiero, Webbia 42: 168 (1988). — Parentage: Ophrys holosericea subsp. apulica × Ophrys sphegodes subsp. atrata.
Ophrys × arachnitiformis nothosubsp. gelana (H.Baumann & Künkele) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × gelana H.Baumann & Künkele, Mitt. Arbeitskreis Heimische Orchid. Baden-Württemberg 18: 492 (1986). — Parentage: Ophrys holosericea subsp. oxyrrhynchos × Ophrys sphegodes subsp. atrata.
Ophrys × arachnitiformis nothosubsp. ingaranensis (Soca) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × ingaranensis Soca, Monde Pl. 91: 18 (1997). — Parentage: Ophrys holosericea subsp. holosericea × Ophrys holosericea subsp. parvimaculata × Ophrys sphegodes.
Ophrys × arachnitiformis nothosubsp. kelleri (Godfery) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × kelleri Godfery, J. Bot. 53: 121 (1915). — Parentage: Ophrys holosericea × Ophrys sphegodes subsp. atrata × Ophrys sphegodes subsp. sphegodes.
Ophrys × arachnitiformis nothosubsp. larissae (Benigni, Barigelli & Petroselli) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × larissae Benigni, Barigelli & Petroselli, GIROS Orch. Spont. Eur. 65: 5 (2022). — Parentage: Ophrys holosericea subsp. apulica × Ophrys sphegodes subsp. araneola.
Ophrys × arachnitiformis nothosubsp. mangiagessoi (Cataldo & S.A.Giardina) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × mangiagessoi Cataldo & S.A.Giardina, J. Eur. Orch. 43: 792 (2011). — Parentage: Ophrys holosericea subsp. biancae × Ophrys sphegodes subsp. sphegodes.
Ophrys × arachnitiformis nothosubsp. marianoi (Licheri & Napoli) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × marianoi Licheri & Napoli, GIROS Orch. Spont. Eur. 68(1): 87 (2025). — Parentage: Ophrys holosericea subsp. holosericea × Ophrys sphegodes subsp. atrata.
Ophrys × arachnitiformis nothosubsp. martorinae (Galesi) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × martorinae Galesi, J. Eur. Orch. 31: 313 (1999). — Parentage: Ophrys holosericea subsp. biancae × Ophrys sphegodes subsp. atrata.
Ophrys × arachnitiformis nothosubsp. montis-lorae (Giotta & Piccitto) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × montis-lorae Giotta & Piccitto, J. Eur. Orch. 44: 761 (2012). — Parentage: Ophrys holosericea subsp. chestermanii × Ophrys sphegodes subsp. atrata.
Ophrys × arachnitiformis nothosubsp. pacis (Benigni, Mandozzi & Stortoni) Z.H.Feng, comb. nov. — Basionym: Ophrys × aschersonii nothosubsp. pacis Benigni, Mandozzi & Stortoni, GIROS Orch. Spont. Eur. 62: 352 (2019). — Parentage: Ophrys holosericea subsp. gracilis × Ophrys sphegodes subsp. sphegodes.
Ophrys × arachnitiformis nothosubsp. pulchra (E.G.Camus) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × pulchra E.G.Camus, Bull. Soc. Bot. France 38: 43 (1891). — Parentage: Ophrys holosericea × Ophrys sphegodes subsp. araneola.
Ophrys × arachnitiformis nothosubsp. rasettii (W.Rossi & Minut.) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × rasettii W.Rossi & Minut., Webbia 43: 351 (1989). — Parentage: Ophrys holosericea subsp. lacaitae × Ophrys sphegodes.
Ophrys × arachnitiformis nothosubsp. reinhardiana (H.Baumann & Künkele) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × reinhardiana H.Baumann & Künkele, Mitt. Arbeitskreis Heimische Orchid. Baden-Württemberg 16: 650 (1984). — Parentage: Ophrys holosericea subsp. oxyrrhynchos × Ophrys sphegodes.
Ophrys × arachnitiformis nothosubsp. sardoa (Licheri) Z.H.Feng, comb. nov. — Basionym: Ophrys × aschersonii nothosubsp. sardoa Licheri, GIROS Orch. Spont. Eur. 63: 448 (2020). — Parentage: Ophrys holosericea subsp. chestermanii × Ophrys sphegodes subsp. sphegodes.
Ophrys × arachnitiformis nothosubsp. sanctimarci (Perilli) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × sanctimarci Perilli, GIROS Orch. Spont. Eur. 58: 311 (2015). — Parentage: Ophrys holosericea subsp. apulica × Ophrys sphegodes subsp. sphegodes.
Ophrys × arachnitiformis nothosubsp. turcoi (Silletti & Medagli) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × turcoi Silletti & Medagli, Quad. Silvae 1: 60 (2015). — Parentage: Ophrys holosericea subsp. apulica × Ophrys holosericea subsp. holosericea × Ophrys sphegodes subsp. sphegodes.
Ophrys × arachnitiformis nothosubsp. turris-veneris (S.Serafino, Chetta & Mascia) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × turris-veneris S.Serafino, Chetta & Mascia, GIROS Orch. Spont. Eur. 65: 302 (2022). — Parentage: Ophrys holosericea subsp. candica × Ophrys sphegodes.
Ophrys × arachnitiformis nothosubsp. velathrii (Romolini & Soca) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × velathrii Romolini & Soca, J. Eur. Orch. 56: 171 (2024). — Parentage: Ophrys holosericea × Ophrys sphegodes subsp. passionis.
Ophrys × benoitiana nothosubsp. goelziana (H.Baumann & Künkele) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × goelziana H.Baumann & Künkele, Mitt. Arbeitskreis Heimische Orchid. Baden-Württemberg 16: 650 (1984). — Parentage: Ophrys lunulata × Ophrys sphegodes subsp. sphegodes.
Ophrys × benoitiana nothosubsp. trullana (H.Baumann & Künkele) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × trullana H.Baumann & Künkele, Mitt. Arbeitskreis Heimische Orchid. Baden-Württemberg 18: 472 (1986). — Parentage: Ophrys lunulata × Ophrys sphegodes subsp. passionis.
Ophrys × borgersiae nothosubsp. mulierum (H.Baumann & Künkele) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × mulierum H.Baumann & Künkele, Mitt. Arbeitskreis Heimische Orchid. Baden-Württemberg 18: 508 (1986). — Parentage: Ophrys reinholdii × Ophrys sphegodes subsp. taurica.
Ophrys × brigittae nothosubsp. bayeri (H.Baumann) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × bayeri H.Baumann, Beih. Veröff. Naturschutz Landschaftspflege Baden-Württemberg 19: 142 (1981). — Parentage: Ophrys fusca subsp. fusca × Ophrys omegaifera subsp. fleischmannii.
Ophrys × brigittae nothosubsp. kurzeorum (H.Baumann) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × kurzeorum H.Baumann, Beih. Veröff. Naturschutz Landschaftspflege Baden-Württemberg 19: 142 (1981). — Parentage: Ophrys fusca subsp. durieui × Ophrys omegaifera subsp. dyris.
Ophrys × brigittae nothosubsp. pauliana (C.Alibertis & A.Alibertis) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × pauliana C.Alibertis & A.Alibertis, Orchidophile (Deuil-la-Barre) 86: 81 (1989). — Parentage: Ophrys fusca subsp. fusca × Ophrys omegaifera subsp. fleischmannii × Ophrys omegaifera subsp. omegaifera.
Ophrys × campolati nothosubsp. lupiae (O.Danesch & E.Danesch) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × lupiae O.Danesch & E.Danesch, Orchid. Eur. Ophrys-Hybr.: 228 (1972). — Parentage: Ophrys bertolonii × Ophrys sphegodes subsp. atrata × Ophrys tenthredinifera.
Ophrys × campolati nothosubsp. varolae (O.Danesch & E.Danesch) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × varolae Lumare, Medagli & Anelli, GIROS Orch. Spont. Eur. 63(1): 185 (2020). — Parentage: Ophrys bertolonii × Ophrys sphegodes subsp. passionis × Ophrys tenthredinifera.
Ophrys × chobautii nothosubsp. mirtiae (G.Kretzschmar & H.Kretzschmar) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × mirtiae G.Kretzschmar & H.Kretzschmar, Ber. Arbeitskreis. Heimische Orchid. 15: 53 (1998). — Parentage: Ophrys lutea subsp. galilaea × Ophrys speculum.
Ophrys × clapensis nothosubsp. domitia (Del Prete) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × domitia Del Prete, Atti Soc. Tosc. Sci. Nat. Pisa, Mem. 85: 270 (1979). — Parentage: Ophrys bombyliflora × Ophrys lutea subsp. murbeckii.
Ophrys × cranbrookeana nothosubsp. lewinii (Soca) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × lewinii Soca, Monde Pl. 91: 14 (1997). — Parentage: Ophrys holosericea subsp. holosericea × Ophrys scolopax subsp. cornuta × Ophrys sphegodes.
Ophrys × cranbrookeana nothosubsp. sutiliae (Robles, J.C.Huertas & M.Becerra) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × sutiliae Robles, J.C.Huertas & M.Becerra, Acta Bot. Malac. 49: 228 (2024). — Parentage: Ophrys holosericea × Ophrys scolopax subsp. apiformis × Ophrys sphegodes.
Ophrys × cugniensis nothosubsp. anxatina (Congedo) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × anxatina Congedo, GIROS Notiz. 36: 34 (2007). — Parentage: Ophrys bertolonii × Ophrys lutea subsp. galilaea.
Ophrys × daneschiana nothosubsp. poisneliae (Menos) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × poisneliae Menos, Orchidophile (Deuil-la-Barre) 143: 197 (2000). — Parentage: Ophrys bertolonii × Ophrys insectifera subsp. aymoninii × Ophrys sphegodes.
Ophrys × eliasii nothosubsp. sanctae-sofiae (M.P.Grasso & Liverani) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × sanctae-sofiae M.P.Grasso & Liverani, Orchidophile (Deuil-la-Barre) 109: 204 (1993). — Parentage: Ophrys fusca subsp. iricolor × Ophrys speculum.
Ophrys × epidavrensis nothosubsp. boscoquartensis (O.Danesch & E.Danesch) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × boscoquartensis O.Danesch & E.Danesch, Orchidee (Hamburg) 21: 361 (1970). — Parentage: Ophrys argolica subsp. biscutella × Ophrys sphegodes.
Ophrys × epidavrensis nothosubsp. castellettae (Benigni, Barigelli & Petroselli) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × castellettae Benigni, Barigelli & Petroselli, GIROS Orch. Spont. Eur. 63: 305 (2020). — Parentage: Ophrys argolica subsp. crabronifera × Ophrys sphegodes subsp. araneola.
Ophrys × epidavrensis nothosubsp. cornelli (O.Danesch & E.Danesch) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × cornelli O.Danesch & E.Danesch, Orchid. Eur. Ophrys-Hybr.: 224 (1972). — Parentage: Ophrys argolica subsp. biscutella × Ophrys sphegodes subsp. sipontensis.
Ophrys × epidavrensis nothosubsp. difesae (Perilli) Z.H.Feng, comb. nov. — Basionym: Ophrys × boscoquartensis nothosubsp. difesae Perilli, GIROS Orch. Spont. Eur. 59: 171 (2016). — Parentage: Ophrys argolica subsp. biscutella × Ophrys sphegodes subsp. passionis.
Ophrys × epidavrensis nothosubsp. floreannae (Licheri & Pani) Z.H.Feng, comb. nov. — Basionym: Ophrys × giardinae nothosubsp. floreannae Licheri & Pani, GIROS Orch. Spont. Eur. 62: 408 (2019). — Parentage: Ophrys argolica subsp. crabronifera × Ophrys sphegodes subsp. passionis × Ophrys sphegodes subsp. sphegodes.
Ophrys × epidavrensis nothosubsp. fluminensis (Licheri & A.S.Loi) Z.H.Feng, comb. nov. — Basionym: Ophrys × nettii nothosubsp. fluminensis Licheri & A.S.Loi, GIROS Orch. Spont. Eur. 62: 196 (2019). — Parentage: Ophrys argolica subsp. crabronifera × Ophrys sphegodes subsp. atrata × Ophrys sphegodes subsp. sphegodes.
Ophrys × epidavrensis nothosubsp. kingsleyana (Klaver & D.Rossi) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × kingsleyana Klaver & D.Rossi, GIROS Orch. Spont. Eur. 64: 51 (2021). — Parentage: Ophrys argolica subsp. crabronifera × Ophrys sphegodes subsp. passionis.
Ophrys × epidavrensis nothosubsp. peloponnesiaca (O.Danesch & E.Danesch) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × peloponnesiaca O.Danesch & E.Danesch, Orchid. Eur. Ophrys-Hybr.: 222 (1972). — Parentage: Ophrys argolica subsp. argolica × Ophrys sphegodes subsp. taurica.
Ophrys × epidavrensis nothosubsp. rabautei (Soca) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × rabautei Soca, Monde Pl. 90: 9 (1995). — Parentage: Ophrys argolica subsp. crabronifera × Ophrys sphegodes subsp. araneola.
Ophrys × epidavrensis nothosubsp. sundermanniana (O.Danesch & E.Danesch) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × sundermanniana O.Danesch & E.Danesch, Orchid. Eur. Ophrys-Hybr.: 223 (1972). — Parentage: Ophrys argolica subsp. argolica × Ophrys sphegodes subsp. spruneri.
Ophrys × feldwegiana nothosubsp. walravensiorum (P.Delforge) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × walravensiorum P.Delforge, Naturalistes Belges 75: 245 (1994). — Parentage: Ophrys ferrum-equinum subsp. gottfriediana × Ophrys tenthredinifera.
Ophrys × ferruginea nothosubsp. marsianii (Galesi) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × marsianii Galesi, J. Eur. Orch. 27: 267 (1995). — Parentage: Ophrys fusca subsp. fusca × Ophrys holosericea subsp. oxyrrhynchos.
Ophrys × flavicans nothosubsp. alberobellensis (H.Baumann & Künkele) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × alberobellensis H.Baumann & Künkele, Mitt. Arbeitskreis Heimische Orchid. Baden-Württemberg 18: 496 (1986). — Parentage: Ophrys bertolonii × Ophrys sphegodes subsp. atrata × Ophrys sphegodes subsp. sphegodes.
Ophrys × flavicans nothosubsp. barlae (E.G.Camus) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × barlae E.G.Camus, J. Bot. (Morot) 7: 159 (1893). — Parentage: Ophrys bertolonii × Ophrys sphegodes subsp. atrata.
Ophrys × flavicans nothosubsp. cantalupoi (Soca) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × cantalupoi Soca, Caesiana 19: 5 (2015). — Parentage: Ophrys bertolonii × Ophrys sphegodes subsp. passionis × Ophrys sphegodes subsp. sphegodes.
Ophrys × flavicans nothosubsp. grottagliensis (P.Delforge & C.Delforge) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × grottagliensis P.Delforge & C.Delforge, Naturalistes Belges 67: 161 (1986). — Parentage: Ophrys bertolonii × Ophrys sphegodes subsp. passionis.
Ophrys × flavicans nothosubsp. neowalteri (A.Camus) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × neowalteri A.Camus, Bull. Mus. Natl. Hist. Nat. 33: 536 (1927). — Parentage: Ophrys bertolonii × Ophrys sphegodes subsp. araneola.
Ophrys × flavicans nothosubsp. pescolusae (Gennaio & Medagli) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × pescolusae Gennaio & Medagli, GIROS Notiz. 54: 22 (2013). — Parentage: Ophrys bertolonii × Ophrys sphegodes subsp. atrata × Ophrys sphegodes subsp. passionis.
Ophrys fusca nothosubsp. dessartiana (P.Delforge) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × dessartiana P.Delforge, Naturalistes Belges 78: 180 (1997). — Parentage: Ophrys fusca subsp. fusca × Ophrys fusca subsp. iricolor.
Ophrys fusca nothosubsp. joannae (Maire) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × joannae Maire, Bull. Soc. Hist. Nat. Afrique N. 12: 49 (1921). — Parentage: Ophrys fusca subsp. durieui × Ophrys fusca subsp. iricolor.
Ophrys fusca nothosubsp. peraiolae (G.Foelsche, W.Foelsche, M.Gerbaud & O.Gerbaud) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × peraiolae G.Foelsche, W.Foelsche, M.Gerbaud & O.Gerbaud, J. Eur. Orch. 32: 421 (2000). — Parentage: Ophrys fusca subsp. bilunulata × Ophrys fusca subsp. funerea.
Ophrys fusca nothosubsp. rafinesquei (Soca) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × rafinesquei Soca, Caesiana 47: 44 (2017). — Parentage: Ophrys fusca subsp. fusca × Ophrys fusca subsp. pallida.
Ophrys × gauthieri nothosubsp. cesinensis (Medagli, D'Emerico & Ruggiero) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × cesinensis Medagli, D'Emerico & Ruggiero, Orchidophile (Deuil-la-Barre) 85: 30 (1989). — Parentage: Ophrys fusca × Ophrys lutea subsp. lutea × Ophrys lutea subsp. galilaea.
Ophrys × gauthieri nothosubsp. heracleotica (Gavalas, Kreutz & Z.Antonop.) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × heracleotica Gavalas, Kreutz & Z.Antonop., Ber. Arbeitskreis. Heimische Orchid. 27: 272 (2011). — Parentage: Ophrys fusca subsp. fusca × Ophrys lutea subsp. galilaea.
Ophrys × gauthieri nothosubsp. paringianensis (Licheri & Napoli) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × paringianensis Licheri & Napoli, GIROS Orch. Spont. Eur. 66: 94 (2023). — Parentage: Ophrys fusca subsp. iricolor × Ophrys lutea subsp. galilaea.
Ophrys × grampinii nothosubsp. alibertiana (C.Alibertis & A.Alibertis) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × alibertiana C.Alibertis & A.Alibertis, Orchidophile (Deuil-la-Barre) 86: 83 (1989). — Parentage: Ophrys sphegodes subsp. spruneri × Ophrys tenthredinifera.
Ophrys × grampinii nothosubsp. caballeroi (C.E.Hermos.) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × caballeroi C.E.Hermos., Estud. Mus. Cienc. Nat. Álava 16: 54 (2001). — Parentage: Ophrys sphegodes subsp. aveyronensis × Ophrys tenthredinifera.
Ophrys × grampinii nothosubsp. manfredoniae (O.Danesch & E.Danesch) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × manfredoniae O.Danesch & E.Danesch, Orchid. Eur. Ophrys-Hybr.: 228 (1972). — Parentage: Ophrys sphegodes subsp. atrata × Ophrys tenthredinifera.
Ophrys × grampinii nothosubsp. robatschii (Benito) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × robatschii Benito, Fl. Montiber. 80: 74 (2021). — Parentage: Ophrys sphegodes subsp. araneola × Ophrys tenthredinifera.
Ophrys × grampinii nothosubsp. surdi (O.Danesch & E.Danesch) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × surdi O.Danesch & E.Danesch, Orchid. Eur. Ophrys-Hybr.: 229 (1972). — Parentage: Ophrys sphegodes subsp. passionis × Ophrys tenthredinifera.
Ophrys × gumprechtii nothosubsp. chettae (Perilli & Biagioli) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × chettae Perilli & Biagioli, GIROS Orch. Spont. Eur. 63: 503 (2020). — Parentage: Ophrys bertolonii × Ophrys holosericea subsp. lacaitae.
Ophrys × gumprechtii nothosubsp. lorenzoi (Bisceglia) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × lorenzoi Bisceglia, GIROS Orch. Spont. Eur. 66: 185 (2023). — Parentage: Ophrys bertolonii × Ophrys holosericea subsp. gracilis.
Ophrys × gumprechtii nothosubsp. nardonei (Lumare, Medagli, Dura & Anelli) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × nardonei Lumare, Medagli, Dura & Anelli, GIROS Orch. Spont. Eur. 62: 287 (2019). — Parentage: Ophrys bertolonii × Ophrys holosericea subsp. apulica × Ophrys holosericea subsp. parvimaculata.
Ophrys × gumprechtii nothosubsp. passalacquae (Silletti & Medagli) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × passalacquae Silletti & Medagli, Quad. Silvae 1: 64 (2015). — Parentage: Ophrys bertolonii × Ophrys holosericea subsp. oxyrrhynchos.
Ophrys holosericea nothosubsp. benignii (Angradi, Simonc., Bocchini & Barch.) Z.H.Feng, comb. nov. — Basionym: Ophrys × benignii Angradi, Simonc., Bocchini & Barch., GIROS Orch. Spont. Eur. 68(1): 183 (2025). — Parentage: Ophrys holosericea subsp. gracilis × Ophrys holosericea subsp. passionis.
Ophrys holosericea nothosubsp. bibbiensis (Soca) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × bibbiensis Soca, Monde Pl. 91: 21 (1997). — Parentage: Ophrys holosericea subsp. biancae × Ophrys holosericea subsp. lacaitae.
Ophrys holosericea nothosubsp. caliandri (O.Danesch & E.Danesch) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × caliandri O.Danesch & E.Danesch, Orchid. Eur. Ophrys-Hybr.: 222 (1972). — Parentage: Ophrys holosericea subsp. apulica × Ophrys holosericea subsp. oxyrrhynchos.
Ophrys holosericea nothosubsp. campoi (Soca) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × campoi Soca, Monde Pl. 95: 13 (2000). — Parentage: Ophrys holosericea subsp. lacaitae × Ophrys holosericea subsp. oxyrrhynchos.
Ophrys holosericea nothosubsp. delargeana (P.Delforge) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × delargeana P.Delforge, Naturalistes Belges 69: 44 (1988). — Parentage: Ophrys holosericea subsp. holosericea × Ophrys holosericea subsp. lacaitae.
Ophrys holosericea nothosubsp. fajarama (Perilli) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × fajarama Perilli, GIROS Orch. Spont. Eur. 58: 313 (2015). — Parentage: Ophrys holosericea subsp. gracilis × Ophrys holosericea subsp. lacaitae.
Ophrys holosericea nothosubsp. mandozzii (Benigni, Barigelli, Petroselli & Bocchini) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × mandozzii Benigni, Barigelli, Petroselli & Bocchini, GIROS Orch. Spont. Eur. 68(1): 177 (2025). — Parentage: Ophrys holosericea subsp. gracilis × Ophrys holosericea subsp. holosericea.
Ophrys holosericea nothosubsp. necropolitana (H.Baumann & Künkele) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × necropolitana H.Baumann & Künkele, Mitt. Arbeitskreis Heimische Orchid. Baden-Württemberg 18: 452 (1986). — Parentage: Ophrys holosericea subsp. biancae × Ophrys holosericea subsp. oxyrrhynchos.
Ophrys holosericea nothosubsp. nociana (H.Baumann & Künkele) Z.H.Feng, comb. nov. — Basionym: Ophrys × cosentiana nothosubsp. nociana H.Baumann & Künkele, Mitt. Arbeitskreis Heimische Orchid. Baden-Württemberg 18: 407 (1986). — Parentage: Ophrys holosericea subsp. apulica × Ophrys holosericea subsp. parvimaculata.
Ophrys holosericea nothosubsp. pescocanalei (Soca) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × pescocanalei Soca, J. Eur. Orch. 46: 671 (2014). — Parentage: Ophrys holosericea subsp. gracilis × Ophrys holosericea subsp. holosericea. — Heterotypic synonym: Ophrys × barigellii Benigni, Petroselli, Mandozzi, Barch. & Bocchini, Micol. Marche, N.S. 9(1): 100 (2025), syn. nov.
Ophrys holosericea nothosubsp. pugliana (H.Baumann & Künkele) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × pugliana H.Baumann & Künkele, Mitt. Arbeitskreis Heimische Orchid. Baden-Württemberg 18: 481 (1986). — Parentage: Ophrys holosericea subsp. holosericea × Ophrys holosericea subsp. oxyrrhynchos.
Ophrys holosericea nothosubsp. renatafonteae (Gennaio, M.Gargiulo, Medagli & Chetta) Z.H.Feng, comb. nov. — Basionym: Ophrys × sivana nothosubsp. renatafonteae Gennaio, M.Gargiulo, Medagli & Chetta, GIROS Orch. Spont. Eur. 59: 417 (2016). — Parentage: Ophrys holosericea subsp. candica × Ophrys holosericea subsp. parvimaculata.
Ophrys holosericea nothosubsp. sivana (H.Baumann & Künkele) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × sivana H.Baumann & Künkele, Mitt. Arbeitskreis Heimische Orchid. Baden-Württemberg 18: 446 (1986). — Parentage: Ophrys holosericea subsp. candica × Ophrys holosericea subsp. holosericea.
Ophrys holosericea nothosubsp. staraleana (D'Alonzo & Perilli) Z.H.Feng, comb. nov. — Basionym: Ophrys × delargeana nothosubsp. staraleana D'Alonzo & Perilli, GIROS Orch. Spont. Eur. 59: 64 (2016). — Parentage: Ophrys holosericea subsp. apulica × Ophrys holosericea subsp. lacaitae.
Ophrys holosericea nothosubsp. valdevariabilis (O.Danesch & E.Danesch) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × valdevariabilis O.Danesch & E.Danesch, Orchid. Eur. Ophrys-Hybr.: 221 (1972). — Parentage: Ophrys holosericea subsp. apulica × Ophrys holosericea subsp. candica.
Ophrys holosericea nothosubsp. vilanensis (Angradi, Simonc. & Benigni) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × vilanensis Angradi, Simonc. & Benigni, GIROS Orch. Spont. Eur. 68(1): 190 (2025). — Parentage: Ophrys holosericea subsp. apulica × Ophrys holosericea subsp. gracilis.
Ophrys × inceburumensis nothosubsp. deheyniana (P.Delforge) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × deheyniana P.Delforge, Naturalistes Belges 78: 180 (1997). — Parentage: Ophrys bornmuelleri × Ophrys scolopax subsp. scolopax.
Ophrys × laconensis nothosubsp. alariensis (S.Serafino) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × alariensis S.Serafino, GIROS Orch. Spont. Eur. 67(2): 179 (2024). — Parentage: Ophrys holosericea subsp. candica × Ophrys sphegodes subsp. passionis × Ophrys tenthredinifera.
Ophrys × laconensis nothosubsp. lattanziae (Lumare, Medagli & Anelli) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × lattanziae Lumare, Medagli & Anelli, GIROS Orch. Spont. Eur. 63(1): 172 (2020). — Parentage: Ophrys holosericea subsp. apulica × Ophrys sphegodes subsp. atrata × Ophrys tenthredinifera.
Ophrys × laconensis nothosubsp. sancti-ligorii (S.Serafino) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × sancti-ligorii S.Serafino, GIROS Orch. Spont. Eur. 65: 292 (2022). — Parentage: Ophrys holosericea subsp. candica × Ophrys sphegodes subsp. atrata × Ophrys tenthredinifera.
Ophrys × luizetii nothosubsp. corvey-bironii (J.-M.Lewin) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × corvey-bironii J.-M.Lewin, Orchidophile (Deuil-la-Barre) 140: 18 (2000). — Parentage: Ophrys apifera × Ophrys sphegodes subsp. aveyronensis.
Ophrys × luizetii nothosubsp. sandrae (Sardu) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × sandrae Sardu, GIROS Orch. Spont. Eur. 61: 357 (2018). — Parentage: Ophrys apifera × Ophrys sphegodes subsp. atrata.
Ophrys × macchiatii nothosubsp. armentariae (Ferrández, Benito & C.E.Hermos.) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × armentariae Ferrández, Benito & C.E.Hermos., Fl. Montiber. 65: 36 (2016). — Parentage: Ophrys speculum × Ophrys sphegodes subsp. passionis.
Ophrys × macchiatii nothosubsp. barsellae (Romolini & Soca) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × barsellae Romolini & Soca, J. Eur. Orch. 56: 185 (2024). — Parentage: Ophrys speculum × Ophrys sphegodes subsp. araneola. — Heterotypic synonym: Ophrys × moscardiniorum Battagl., Romolini & Soca, J. Eur. Orch. 56: 60 (2024), syn. nov.
Ophrys × macchiatii nothosubsp. garnierae (P.Delforge) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × garnierae P.Delforge, Orchidophile (Deuil-la-Barre) 59: 497 (1983). — Parentage: Ophrys speculum × Ophrys sphegodes subsp. spruneri.
Ophrys × macchiatii nothosubsp. lydia (H.Baumann & Künkele) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × lydia H.Baumann & Künkele, Mitt. Arbeitskreis Heimische Orchid. Baden-Württemberg 18: 511 (1986). — Parentage: Ophrys speculum subsp. regis-ferdinandii × Ophrys sphegodes subsp. taurica.
Ophrys × maremmae nothosubsp. arciensis (Bartolo & Pulv.) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × arciensis Bartolo & Pulv., J. Eur. Orch. 30: 647 (1998). — Parentage: Ophrys holosericea subsp. oxyrrhynchos × Ophrys tenthredinifera.
Ophrys × maremmae nothosubsp. hydruntensis (O.Danesch & E.Danesch) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × hydruntensis O.Danesch & E.Danesch, Orchid. Eur. Ophrys-Hybr.: 221 (1972). — Parentage: Ophrys holosericea subsp. apulica × Ophrys holosericea subsp. candica × Ophrys tenthredinifera.
Ophrys × maremmae nothosubsp. romanoi (D'Alonzo & Perilli) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × romanoi D'Alonzo & Perilli, GIROS Orch. Spont. Eur. 62: 171 (2019). — Parentage: Ophrys holosericea subsp. lacaitae × Ophrys tenthredinifera.
Ophrys × maremmae nothosubsp. salentina (O.Danesch & E.Danesch) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × salentina O.Danesch & E.Danesch, Orchid. Eur. Ophrys-Hybr.: 222 (1972). — Parentage: Ophrys holosericea subsp. apulica × Ophrys tenthredinifera.
Ophrys × marmarensis nothosubsp. liebischiana (Kohlmüller) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × liebischiana Kohlmüller, Mitt. Arbeitskreis Heimische Orchid. Baden-Württemberg 22: 816 (1990). — Parentage: Ophrys holosericea subsp. grandiflora × Ophrys umbilicata subsp. lapethica.
Ophrys × marmarensis nothosubsp. willingiorum (H.Baumann & Künkele) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × willingiorum H.Baumann & Künkele, Mitt. Arbeitskreis Heimische Orchid. Baden-Württemberg 18: 501 (1986). — Parentage: Ophrys holosericea subsp. grandiflora × Ophrys umbilicata.
Ophrys × methonensis nothosubsp. simica (Keitel & Remm) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × simica Keitel & Remm, Mitt. Arbeitskreis Heimische Orchid. Baden-Württemberg 23: 90 (1991). — Parentage: Ophrys argolica subsp. lucis × Ophrys tenthredinifera.
Ophrys × methonensis nothosubsp. tuscanica (H.Baumann & Künkele) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × tuscanica H.Baumann & Künkele, Mitt. Arbeitskreis Heimische Orchid. Baden-Württemberg 18: 450 (1986). — Parentage: Ophrys argolica subsp. crabronifera × Ophrys tenthredinifera.
Ophrys × minuticauda nothosubsp. pompelonensis (Robles) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × pompelonensis Robles, M.Becerra & A.Becerra, Acta Bot. Malac. 46e10589: (2021). — Parentage: Ophrys apifera × Ophrys scolopax subsp. apiformis.
Ophrys × moreana nothosubsp. telchinensis (Riech.) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × telchinensis Riech., Ber. Arbeitskreis. Heimische Orchid. 14: 67 (1997). — Parentage: Ophrys argolica subsp. lucis × Ophrys ferrum-equinum.
Ophrys × neocamusii nothosubsp. santeramana (Silletti & Medagli) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × santeramana Silletti & Medagli, Quad. Silvae 1: 58 (2015). — Parentage: Ophrys bertolonii × Ophrys holosericea × Ophrys sphegodes subsp. atrata × Ophrys sphegodes subsp. sphegodes.
Ophrys × neocamusii nothosubsp. hilleri (Cristaudo, Galesi, R.Lorenz & Zelesny) Z.H.Feng, comb. nov. — Basionym: Ophrys × valparmensis nothosubsp. hilleri Cristaudo, Galesi, R.Lorenz & Zelesny, J. Eur. Orch. 42: 190 (2010). — Parentage: Ophrys bertolonii × Ophrys holosericea subsp. apulica × Ophrys sphegodes.
Ophrys × neoruppertii nothosubsp. littoralis (J.Baláz & M.Baláz) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × littoralis J.Baláz & M.Baláz, Orchidee (Hamburg) 40: 108 (1989). — Parentage: Ophrys bertolonii × Ophrys scolopax subsp. cornuta.
Ophrys × nouletii nothosubsp. aghemanii (Renz) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × aghemanii Renz, Fl. Iranica 126: 86 (1978). — Parentage: Ophrys scolopax subsp. cornuta × Ophrys sphegodes subsp. taurica.
Ophrys × nouletii nothosubsp. calenae (O.Danesch & E.Danesch) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys philippii nm. calenae O.Danesch & E.Danesch, Orchid. Eur. Ophrys-Hybriden 228, 97 (1972). — Parentage: Ophrys scolopax subsp. cornuta × Ophrys sphegodes subsp. sphegodes.
Ophrys × nouletii nothosubsp. bernardii (Looken) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × bernardii Looken, Orchidophile (Deuil-la-Barre) 75: 1211 (1987). — Parentage: Ophrys scolopax subsp. scolopax × Ophrys sphegodes subsp. aveyronensis.
Ophrys × nouletii nothosubsp. disabatoi (D'Alonzo & Perilli) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × disabatoi D'Alonzo & Perilli, GIROS Orch. Spont. Eur. 61: 6 (2018). — Parentage: Ophrys scolopax subsp. cornuta × Ophrys sphegodes subsp. passionis.
Ophrys × nouletii nothosubsp. dodonica (B.Willing & E.Willing) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × dodonica B.Willing & E.Willing, Orchidophile (Deuil-la-Barre) 72: 110 (1986). — Parentage: Ophrys scolopax subsp. cornuta × Ophrys sphegodes subsp. helenae.
Ophrys × nouletii nothosubsp. duvigneaudiana (P.Delforge & C.Delforge) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × duvigneaudiana P.Delforge & C.Delforge, Orchidophile (Deuil-la-Barre) 78: 1364 (1987). — Parentage: Ophrys scolopax subsp. scolopax × Ophrys sphegodes subsp. araneola.
Ophrys × nouletii nothosubsp. kiparissensis (B.Willing & E.Willing) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × kiparissensis B.Willing & E.Willing, Mitt. Arbeitskreis Heimische Orchid. Baden-Württemberg 10: 134 (1978). — Parentage: Ophrys scolopax subsp. cornuta × Ophrys sphegodes subsp. spruneri.
Ophrys × nouletii nothosubsp. llenasii (Sennen ex E.G.Camus) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × llenasii Sennen ex E.G.Camus, Iconogr. Orchid. Europe, Texte: 356 (1928). — Parentage: Ophrys scolopax subsp. scolopax × Ophrys sphegodes subsp. atrata.
Ophrys × nouletii nothosubsp. muellneri (H.Fleischm.) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × muellneri H.Fleischm., Verh. K. K. Zool.-Bot. Ges. Wien 54: 471 (1904). — Parentage: Ophrys scolopax subsp. cornuta × Ophrys sphegodes subsp. araneola.
Ophrys × nouletii nothosubsp. sooi (A.Fuchs) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × sooi A.Fuchs, Repert. Spec. Nov. Regni Veg. Beih. 51: 131 (1928). — Parentage: Ophrys scolopax subsp. cornuta × Ophrys sphegodes subsp. atrata.
Ophrys × olbiensis nothosubsp. benaventei (Robles & M.Becerra) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × benaventei Robles & M.Becerra, Fl. Montiber. 82: 10 (2022). — Parentage: Ophrys bombyliflora × Ophrys scolopax subsp. apiformis.
Ophrys × olbiensis nothosubsp. lefevreana (P.Delforge) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × lefevreana P.Delforge, Naturalistes Belges 87(Orchid. 19): 219 (2006). — Parentage: Ophrys bombyliflora × Ophrys scolopax subsp. heldreichii.
Ophrys omegaifera nothosubsp. asterusica (C.Alibertis & A.Alibertis) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × asterusica C.Alibertis & A.Alibertis, Orchidophile (Deuil-la-Barre) 86: 81 (1989). — Parentage: Ophrys omegaifera subsp. fleischmannii × Ophrys omegaifera subsp. omegaifera.
Ophrys × paphosiana nothosubsp. denglerensis (Riech) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × denglerensis Riech., Ber. Arbeitskreis. Heimische Orchid. 15: 59 (1998). — Parentage: Ophrys kotschyi × Ophrys umbilicata subsp. lapethica.
Ophrys × peltieri nothosubsp. aranii (Robles, M.Becerra, Astete & P.Barrena) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × aranii Robles, M.Becerra, Astete & P.Barrena, Fl. Montiber. 80: 74 (2021). — Parentage: Ophrys scolopax subsp. apiformis × Ophrys tenthredinifera.
Ophrys × peltieri nothosubsp. lioniana (P.Delforge) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × lioniana P.Delforge, Naturalistes Belges 87(Orchid. 19): 225 (2006). — Parentage: Ophrys scolopax subsp. rhodia × Ophrys tenthredinifera.
Ophrys × peltieri nothosubsp. messeniensis (H.Baumann & Künkele) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × messeniensis H.Baumann & Künkele, Mitt. Arbeitskreis Heimische Orchid. Baden-Württemberg 18: 514 (1986). — Parentage: Ophrys scolopax subsp. cornuta × Ophrys tenthredinifera.
Ophrys × personei nothosubsp. triadensis (G.Kretzschmar & H.Kretzschmar) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × triadensis G.Kretzschmar & H.Kretzschmar, Ber. Arbeitskreis. Heimische Orchid. 15: 52 (1998). — Parentage: Ophrys lutea subsp. galilaea × Ophrys tenthredinifera. — Heterotypic synonym: Ophrys × lumarei Medagli & Baldari, GIROS Orch. Spont. Eur. 68(1): 52 (2025), syn. nov. — Ophrys × mathiae Licheri, Medau & Napoli, GIROS Orch. Spont. Eur. 68(2): 338 (2025), syn. nov.
Ophrys × pseudofusca nothosubsp. boninii (D'Alonzo & Perilli) Z.H.Feng, comb. nov. — Basionym: Ophrys × friderici nothosubsp. boninii D'Alonzo & Perilli, GIROS Orch. Spont. Eur. 63(1): 30 (2020). — Parentage: Ophrys fusca subsp. iricolor × Ophrys sphegodes subsp. sipontensis.
Ophrys × pseudofusca nothosubsp. borakisiana (Albert & Camus) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × borakisiana P.Delforge & C.Delforge, Orchidophile (Deuil-la-Barre) 63: 687 (1984). — Parentage: Ophrys fusca subsp. fusca × Ophrys sphegodes subsp. taurica.
Ophrys × pseudofusca nothosubsp. braun-blanquetiana (Soó) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × braun-blanquetiana Soó, Repert. Spec. Nov. Regni Veg. 26: 280 (1929). — Parentage: Ophrys fusca subsp. fusca × Ophrys sphegodes subsp. atrata.
Ophrys × pseudofusca nothosubsp. capellae-pacis (G.Kretzschmar & H.Kretzschmar) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × capellae-pacis G.Kretzschmar & H.Kretzschmar, Ber. Arbeitskreis. Heimische Orchid. 20: 70 (2003). — Parentage: Ophrys fusca subsp. iricolor × Ophrys sphegodes subsp. spruneri.
Ophrys × pseudofusca nothosubsp. gumprechtiana (Soó) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × gumprechtiana Soó, Acta Bot. Acad. Sci. Hung. 25: 362 (1980). — Parentage: Ophrys fusca subsp. fusca × Ophrys sphegodes subsp. passionis.
Ophrys × pseudofusca nothosubsp. norae (Argiolas) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × norae Argiolas, GIROS Orch. Spont. Eur. 61: 417, 419 (2018). — Parentage: Ophrys fusca subsp. iricolor × Ophrys sphegodes subsp. passionis.
Ophrys × pseudofusca nothosubsp. pano-lefkaron (Bourdon) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × pano-lefkaron Bourdon, Orchidophile (Deuil-la-Barre) 71: 1056 (1986). — Parentage: Ophrys fusca subsp. iricolor × Ophrys sphegodes subsp. taurica.
Ophrys × pseudofusca nothosubsp. pourteiniae (J.M.Mathé) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × pourteiniae J.M.Mathé, Orchidophile (Deuil-la-Barre) 140: 25 (2000). — Parentage: Ophrys fusca subsp. fusca × Ophrys sphegodes subsp. araneola.
Ophrys × pseudofusca nothosubsp. souliei (Soca) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × souliei Soca, Caesiana 24: 21 (2005). — Parentage: Ophrys fusca subsp. fusca × Ophrys sphegodes subsp. aveyronensis.
Ophrys × pseudoquadriloba nothosubsp. ballerinii (D'Alonzo & Perilli) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × ballerinii D'Alonzo & Perilli, GIROS Orch. Spont. Eur. 63(1): 26 (2020). — Parentage: Ophrys lutea subsp. galilaea × Ophrys sphegodes subsp. sipontensis.
Ophrys × pseudoquadriloba nothosubsp. coletteae (D'Alonzo & Perilli) Z.H.Feng, comb. nov. — Basionym: Ophrys × mirandana nothosubsp. coletteae D'Alonzo & Perilli, GIROS Orch. Spont. Eur. 61: 400, 402 (2018). — Parentage: Ophrys lutea subsp. galilaea × Ophrys sphegodes subsp. passionis.
Ophrys × pseudoquadriloba nothosubsp. demangeana (P.Delforge) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × demangeana P.Delforge, Naturalistes Belges 71: 135 (1990). — Parentage: Ophrys lutea subsp. galilaea × Ophrys sphegodes subsp. taurica.
Ophrys × pseudoquadriloba nothosubsp. leguerrierae (P.Delforge) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × leguerrierae P.Delforge, Orchidophile (Deuil-la-Barre) 60: 517 (1984). — Parentage: Ophrys lutea subsp. lutea × Ophrys sphegodes subsp. araneola.
Ophrys × pseudoquadriloba nothosubsp. maschioi (D'Alonzo & Perilli) Z.H.Feng, comb. nov. — Basionym: Ophrys × mirandana nothosubsp. maschioi D'Alonzo & Perilli, GIROS Orch. Spont. Eur. 64: 71 (2021). — Parentage: Ophrys lutea subsp. lutea × Ophrys sphegodes subsp. passionis.
Ophrys × pseudoquadriloba nothosubsp. mattinatellae (Kohlmüller) Z.H.Feng, comb. nov. — Basionym: Ophrys × piscinica nothosubsp. mattinatellae Kohlmüller, Orchidee (Hamburg) 44: 98 (1993). — Parentage: Ophrys lutea subsp. galilaea × Ophrys sphegodes subsp. atrata. — Heterotypic synonym: Ophrys × johannis-patris Licheri & Napoli, GIROS Orch. Spont. Eur. 68(2): 345 (2025), syn. nov.
Ophrys × pseudoquadriloba nothosubsp. piscinica (P.Delforge & C.Delforge) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × piscinica P.Delforge & C.Delforge, Naturalistes Belges 67: 157 (1986). — Parentage: Ophrys lutea subsp. lutea × Ophrys sphegodes subsp. atrata.
Ophrys × pseudoquadriloba nothosubsp. tuchulchae (Antonj & Gransinigh) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × tuchulchae Antonj & Gransinigh, GIROS Orch. Spont. Eur. 68(1): 63 (2025). — Parentage: Ophrys lutea subsp. galilaea × Ophrys sphegodes subsp. araneola.
Ophrys × pseudospeculum nothosubsp. oretana (Robles, A.M.Díaz & Miguel Gómez) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × oretana Robles, A.M.Díaz & Miguel Gómez, Fl. Montiber. 82: 9 (2022). — Parentage: Ophrys lutea × Ophrys scolopax subsp. apiformis.
Ophrys × rainei nothosubsp. daunia (H.Baumann & Künkele) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × daunia H.Baumann & Künkele, Mitt. Arbeitskreis Heimische Orchid. Baden-Württemberg 18: 436 (1986). — Parentage: Ophrys bombyliflora × Ophrys sphegodes subsp. passionis.
Ophrys × rainei nothosubsp. stivii (Romolini & Soca) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × stivii Romolini & Soca, GIROS Orch. Spont. Eur. 58: 157 (2015). — Parentage: Ophrys bombyliflora × Ophrys sphegodes subsp. araneola.
Ophrys × rasbachii nothosubsp. subertana (Battagl., Romolini & Soca) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × subertana Battagl., Romolini & Soca, J. Eur. Orch. 56: 62 (2024). — Parentage: Ophrys argolica subsp. crabronifera × Ophrys speculum.
Ophrys × raynaudii nothosubsp. konyana (Kreutz & Ruedi Peter) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × konyana Kreutz & Ruedi Peter, J. Eur. Orch. 39: 73 (2007). — Parentage: Ophrys argolica subsp. lucis × Ophrys reinholdii subsp. straussii.
Ophrys × rechingeri nothosubsp. demutiana (B.Baumann & H.Baumann) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × demutiana B.Baumann & H.Baumann, Mitt. Arbeitskreis Heimische Orchid. Baden-Württemberg 16: 126 (1984). — Parentage: Ophrys ferrum-equinum subsp. gottfriediana × Ophrys sphegodes subsp. spruneri.
Ophrys × rechingeri nothosubsp. fortosica (B.Willing & E.Willing) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × fortosica B.Willing & E.Willing, Orchidophile (Deuil-la-Barre) 72: 1102 (1986). — Parentage: Ophrys ferrum-equinum × Ophrys sphegodes subsp. helenae.
Ophrys × rechingeri nothosubsp. mastii (P.Delforge) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × mastii P.Delforge, Orchidophile (Deuil-la-Barre) 59: 497 (1983). — Parentage: Ophrys ferrum-equinum × Ophrys sphegodes subsp. spruneri.
Ophrys × resurrecta nothosubsp. biagiolii (Lumare & Medagli) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × biagiolii Lumare & Medagli, GIROS Orch. Spont. Eur. 61: 364, 370 (2018). — Parentage: Ophrys bombyliflora × Ophrys holosericea subsp. candica × Ophrys holosericea subsp. parvimaculata.
Ophrys × resurrecta nothosubsp. martini (O.Danesch & E.Danesch) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × martini O.Danesch & E.Danesch, Orchid. Eur. Ophrys-Hybr.: 227 (1972). — Parentage: Ophrys bombyliflora × Ophrys holosericea subsp. oxyrrhynchos.
Ophrys × resurrecta nothosubsp. medaglii (Turco, Ruggiero, Gennaio & D'Emerico) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × medaglii Turco, Ruggiero, Gennaio & D'Emerico, Thalassia Salentina 34: 27 (2012). — Parentage: Ophrys bombyliflora × Ophrys holosericea subsp. candica.
Ophrys × resurrecta nothosubsp. ozantina (Gennaio) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × ozantina Gennaio, GIROS Notiz. 41: 28 (2009). — Parentage: Ophrys bombyliflora × Ophrys holosericea subsp. parvimaculata.
Ophrys × resurrecta nothosubsp. salapiensis (Serafino, Gennaio, Chetta, M.Gargiulo & Medagli) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × salapiensis Serafino, Gennaio, Chetta, M.Gargiulo & Medagli, GIROS Orch. Spont. Eur. 60: 442 (2017). — Parentage: Ophrys bombyliflora × Ophrys holosericea subsp. apulica × Ophrys holosericea subsp. candica.
Ophrys scolopax nothosubsp. vereeckeniana (P.Delforge) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × vereeckeniana P.Delforge, Naturalistes Belges 87(Orchid. 19): 225 (2006). — Parentage: Ophrys scolopax subsp. heldreichii × Ophrys scolopax subsp. rhodia.
Ophrys speculum nothosubsp. innominata (D.Tyteca & B.Tyteca) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × innominata D.Tyteca & B.Tyteca, Orchidophile (Deuil-la-Barre) 69: 929 (1985). — Parentage: Ophrys speculum subsp. lusitanica × Ophrys speculum subsp. speculum.
Ophrys sphegodes nothosubsp. acina (C.E.Hermos.) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × acina C.E.Hermos., J. Eur. Orch. 31(4): 883 (1999). — Parentage: Ophrys sphegodes subsp. araneola × Ophrys sphegodes subsp. passionis. — Heterotypic synonym: Ophrys × petrosellii Benigni, Barigelli, Mandozzi, Barch. & Bocchini, Micol. Marche, N.S. 9(1): 101 (2025), syn. nov.
Ophrys sphegodes nothosubsp. biancoae (Medagli, D'Emerico & Ruggiero) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × biancoae Medagli, D'Emerico & Ruggiero, Orchidophile (Deuil-la-Barre) 85: 29 (1989). — Parentage: Ophrys sphegodes subsp. passionis × Ophrys sphegodes subsp. sphegodes.
Ophrys sphegodes nothosubsp. burneriana (C.Alibertis & A.Alibertis) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × burneriana C.Alibertis & A.Alibertis, Orchidophile (Deuil-la-Barre) 86: 83 (1989). — Parentage: Ophrys sphegodes subsp. cretensis × Ophrys sphegodes subsp. spruneri.
Ophrys sphegodes nothosubsp. celani (O.Danesch & E.Danesch) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × celani O.Danesch & E.Danesch, Orchid. Eur. Ophrys-Hybr.: 223 (1972). — Parentage: Ophrys sphegodes subsp. atrata × Ophrys sphegodes subsp. passionis.
Ophrys sphegodes nothosubsp. cortesii (A.Camus) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × cortesii A.Camus, Iconogr. Orchid. Europe, Texte: 5 (1921). — Parentage: Ophrys sphegodes subsp. araneola × Ophrys sphegodes subsp. atrata.
Ophrys sphegodes nothosubsp. costei (Looken) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × costei Looken, Orchidophile (Deuil-la-Barre) 86: 84 (1989). — Parentage: Ophrys sphegodes subsp. aveyronensis × Ophrys sphegodes subsp. sphegodes.
Ophrys sphegodes nothosubsp. delvauxiorum (P.Delforge) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × delvauxiorum P.Delforge, Naturalistes Belges 73: 99 (1992). — Parentage: Ophrys sphegodes subsp. helenae × Ophrys sphegodes subsp. sphegodes.
Ophrys sphegodes nothosubsp. evvia (Riech.) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × evvia Riech., Ber. Arbeitskreis. Heimische Orchid. 17: 45 (2000). — Parentage: Ophrys sphegodes subsp. aesculapii × Ophrys sphegodes subsp. taurica.
Ophrys sphegodes nothosubsp. hierapetrae (H.Baumann & Künkele) Z.H.Feng, comb. nov. — Basionym: Ophrys × baumanniana nothosubsp. hierapetrae H.Baumann & Künkele, Mitt. Arbeitskreis Heimische Orchid. Baden-Württemberg 18: 454 (1986). — Parentage: Ophrys sphegodes subsp. cretensis × Ophrys sphegodes subsp. parnassica.
Ophrys sphegodes nothosubsp. ioanninensis (B.Willing & E.Willing) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × ioanninensis B.Willing & E.Willing, Orchidophile (Deuil-la-Barre) 72: 1100 (1986). — Parentage: Ophrys sphegodes subsp. helenae × Ophrys sphegodes subsp. taurica.
Ophrys sphegodes nothosubsp. montana (H.Baumann & Künkele) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × montana H.Baumann & Künkele, Mitt. Arbeitskreis Heimische Orchid. Baden-Württemberg 18: 474 (1986). — Parentage: Ophrys sphegodes subsp. sphegodes × Ophrys sphegodes subsp. taurica.
Ophrys sphegodes nothosubsp. perrinii (Soca) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × perrinii Soca, Monde Pl. 95: 14 (2000). — Parentage: Ophrys sphegodes subsp. passionis × Ophrys sphegodes subsp. provincialis.
Ophrys sphegodes nothosubsp. pseudospruneri (Soó) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × pseudospruneri Soó, Repert. Spec. Nov. Regni Veg. 26: 280 (1929). — Parentage: Ophrys sphegodes subsp. spruneri × Ophrys sphegodes subsp. taurica.
Ophrys sphegodes nothosubsp. sassettae (Romolini & Soca) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × sassettae Romolini & Soca, GIROS Notiz. 38: 17 (2008). — Parentage: Ophrys sphegodes subsp. araneola × Ophrys sphegodes subsp. sphegodes.
Ophrys sphegodes nothosubsp. sieberi (H.Baumann & Künkele) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × sieberi H.Baumann & Künkele, Mitt. Arbeitskreis Heimische Orchid. Baden-Württemberg 18: 453 (1986). — Parentage: Ophrys sphegodes subsp. parnassica × Ophrys sphegodes subsp. taurica.
Ophrys sphegodes nothosubsp. turchinae (Gransinigh & Buono) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × turchinae Gransinigh & Buono, GIROS Notiz. 47: 75 (2011). — Parentage: Ophrys sphegodes subsp. araneola × Ophrys sphegodes subsp. atrata.
Ophrys sphegodes nothosubsp. todaroana (Macch.) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × todaroana Macch., Nuovo Giorn. Bot. Ital. 13: 314 (1881). — Parentage: Ophrys sphegodes subsp. atrata × Ophrys sphegodes subsp. sphegodes.
Ophrys × vicina nothosubsp. anomala (Renz) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × anomala Renz, Repert. Spec. Nov. Regni Veg. 30: 109 (1932). — Parentage: Ophrys holosericea subsp. holosericea × Ophrys scolopax subsp. heldreichii.
Ophrys × vicina nothosubsp. colonnae (D'Alonzo & Perilli) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × colonnae D'Alonzo & Perilli, GIROS Orch. Spont. Eur. 61: 72 (2018). — Parentage: Ophrys holosericea subsp. gracilis × Ophrys scolopax subsp. conradiae.
Ophrys × vicina nothosubsp. deleoi (D'Alonzo & Perilli) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × deleoi D'Alonzo & Perilli, GIROS Orch. Spont. Eur. 63: 328 (2020). — Parentage: Ophrys holosericea subsp. holosericea × Ophrys scolopax subsp. conradiae.
Ophrys × vicina nothosubsp. difesagrandeana (D'Alonzo) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × difesagrandeana D'Alonzo, GIROS Notiz. 57: 32 (2014). — Parentage: Ophrys holosericea subsp. apulica × Ophrys scolopax subsp. conradiae.
Ophrys × vicina nothosubsp. duchateauana (P.Delforge) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × duchateauana P.Delforge, Naturalistes Belges 92: 30 (2011). — Parentage: Ophrys holosericea subsp. candica × Ophrys scolopax subsp. cornuta.
Ophrys × vicina nothosubsp. neozimmermannii (A.Camus) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × neozimmermannii A.Camus, Iconogr. Orchid. Europe, Texte: 357 (1928). — Parentage: Ophrys holosericea subsp. holosericea × Ophrys scolopax subsp. cornuta.
Ophrys × vicina nothosubsp. perspicua (Renz & Taubenheim) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × perspicua Renz & Taubenheim, Notes Roy. Bot. Gard. Edinburgh 41: 276 (1983). — Parentage: Ophrys holosericea subsp. holosericea × Ophrys scolopax subsp. phrygia.
Ophrys × vicina nothosubsp. saskiana (Läpple) Z.H.Feng, comb. nov. — Basionym: Ophrys × marmarensis nothosubsp. saskiana Läpple, Mitt. Arbeitskreis Heimische Orchid. Baden-Württemberg 24: 627 (1992). — Parentage: Ophrys holosericea subsp. holosericea × Ophrys scolopax subsp. rhodia.
Ophrys × vicina nothosubsp. turrica (R.Lorenz & Gembardt) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × turrica R.Lorenz & Gembardt, Mitt. Arbeitskreis Heimische Orchid. Baden-Württemberg 19: 514 (1987). — Parentage: Ophrys holosericea subsp. apulica × Ophrys scolopax subsp. cornuta.
Ophrys × vicina nothosubsp. warwarensis (H.Baumann & Künkele) Z.H.Feng, comb. & stat. nov. — Basionym: Ophrys × warwarensis H.Baumann & Künkele, Mitt. Arbeitskreis Heimische Orchid. Baden-Württemberg 18: 445 (1986). — Parentage: Ophrys holosericea subsp. candica × Ophrys scolopax subsp. heldreichii.
Serapias × meridionalis nothosubsp. pulae (Perko) Z.H.Feng & J.M.H. Shaw, comb. & stat. nov. — Basionym: Serapias × pulae Perko, Ber. Arbeitskreis. Heimische Orchid. 15: 25 (1998). — Parentage: Serapias lingua × Serapias neglecta subsp. apulica.
× Serapicamptis correvonii nothosubsp. rana (Frosch) Z.H.Feng, comb. & stat. nov. — Basionym: × Orchiserapias rana Frosch, Orchidee (Hamburg) 40: 227 (1980). — Parentage: Anacamptis morio subsp. longicornu × Serapias lingua.
Paeoniaceae
Paeonia daurica nothosubsp. dmitrievae (Timukhin & K.U.Aliev) Z.H.Feng, comb. & stat. nov. — Basionym: Paeonia × dmitrievae Timukhin & K.U.Aliev, Turczaninowia 21(3): 51 (2018). — Parentage: Paeonia daurica subsp. coriifolia × Paeonia daurica subsp. macrophylla.
Paeonia daurica nothosubsp. litvinskajae (Mordak, Punina & Timukhin) Z.H.Feng, comb. & stat. nov. — Basionym: Paeonia × litvinskajae Mordak, Punina & Timukhin, Novosti Sist. Vyssh. Rast. 42: 124 (2011). — Parentage: Paeonia daurica subsp. coriifolia × Paeonia daurica subsp. wittmanniana.
Papaveraceae
Papaver pilosum nothosubsp. pichlerianum (Fedde) Z.H.Feng, comb. & stat. nov. — Basionym: Papaver × pichlerianum Fedde, Pflanzenr. (Engler) IV, 104: 358 (1909). — Parentage: Papaver pilosum subsp. pilosum × Papaver pilosum subsp. spicatum.
Papaver pilosum nothosubsp. sieheanum (Fedde) Z.H.Feng, comb. & stat. nov. — Basionym: Papaver × sieheanum Fedde, Pflanzenr. (Engler) IV, 104: 359 (1909). — Parentage: Papaver pilosum × Papaver pilosum subsp. strictum.
Poaceae
Hordeum vulgare nothosubsp. lagunculciforme (Bachteev) Z.H.Feng, comb. & stat. nov. — Basionym: Hordeum spontaneum var. lagunculciforme Bachteev, Bot. Zhurn. (Moscow & Leningrad) 47: 847 (1962). — Parentage: Hordeum vulgare subsp. spontaneum × Hordeum vulgare subsp. vulgare.
Primulaceae
Cyclamen coum nothosubsp. atkinsii (T.Moore) Z.H.Feng, comb. & stat. nov. — Basionym: Cyclamen × atkinsii T.Moore, Gard. Companion Florists' Guide 1: 89 (1852). — Parentage: Cyclamen coum subsp. caucasicum × Cyclamen coum subsp. coum.
Cyclamen purpurascens nothosubsp. marsii (Halda) Z.H.Feng, comb. & stat. nov. — Basionym: Cyclamen × marsii Halda, Skalničky 1973(1): 28 (1973). — Parentage: Cyclamen purpurascens subsp. immaculatum × Cyclamen purpurascens subsp. purpurascens.
Primula × austriaca nothosubsp. gaisbergensis (Pax) Z.H.Feng, comb. & stat. nov. — Basionym: Primula × gaisbergensis Pax, Pflanzenr. (Engler), IV, 237: 64 (1905). — Parentage: Primula veris subsp. canescens × Primula vulgaris.
Primula × austriaca nothosubsp. tristis (Pax) Z.H.Feng, comb. & stat. nov. — Basionym: Primula × tristis Pax, Pflanzenr. (Engler), IV, 237: 65 (1905). — Parentage: Primula veris subsp. veris × Primula vulgaris subsp. rubra.
Primula × austriaca nothosubsp. uzungolensis (Terzioğlu & Coşkunç.) Z.H.Feng, comb. & stat. nov. — Basionym: Primula × uzungolensis Terzioğlu & Coşkunç., Turk. J. Bot. 36(1): 12 (2012). — Parentage: Primula veris subsp. columnae × Primula vulgaris subsp. rubra.
Saxifragaceae
Micranthes nelsoniana nothovar. tolmatchevii (Zhmylev) Z.H.Feng, comb. & stat. nov. — Basionym: Saxifraga × tolmatchevii Zhmylev, Byull. Moskovsk. Obshch. Isp. Prir., Otd. Biol. 100(3): 81 (1995). — Parentage: Micranthes nelsoniana var. nelsoniana × Micranthes nelsoniana var. porsildiana.
3. Conclusion
The current article serves as an overhauled addendum of nomenclature to the existing nothotaxa, these nomenclatural updates are not merely academic exercises; they serve essential practical functions. By validating and standardizing names for overlooked hybrids, this work supports more accurate naming, enhances biodiversity documentation, and provides a reliable reference for conservation prioritization and horticultural use. Furthermore, the integration of these names into major databases will facilitate their adoption in global taxonomic checklists and floristic treatments.
Abbreviations

APG

Angiosperm Phylogeny Group

ICN

International Code of Nomenclature for Algae, Fungi, and Plants

IPNI

International Plant Name Index

POWO

Plants of the World Online

PPG

Pteridophyte Phylogeny Group

Acknowledgments
I am deeply indebted to Dr. Julian M.H. Shaw, from the Royal Horticultural Society, for his insightful discussion on nomenclature of orchids with expertise. I truely appreciate Mr. Kai-Wen Jiang, from South China National Botanical Garden (SCBG), for his valuable comments on leguminous taxonomy with enthusiasm. My thanks go to Ms. Ying-ying Wu for polishing language. I thank colleagues of BAZI for critically reviewing the manuscript before submission and for keen comments and advice from anonymous referee(s) that contribute to the final version of the manuscript.
Author Contributions
Zhen-Hao Feng is the sole author. The author read and approved the final manuscript.
Conflicts of Interest
The author declares no conflicts of interest.
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Cite This Article
  • APA Style

    Feng, Z. (2026). Gathering Pearls from the Lethe: Nomenclatural Novelties for Nothotaxa. Plant, 14(1), 1-23. https://doi.org/10.11648/j.plant.20261401.11

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    Feng, Z. Gathering Pearls from the Lethe: Nomenclatural Novelties for Nothotaxa. Plant. 2026, 14(1), 1-23. doi: 10.11648/j.plant.20261401.11

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    AMA Style

    Feng Z. Gathering Pearls from the Lethe: Nomenclatural Novelties for Nothotaxa. Plant. 2026;14(1):1-23. doi: 10.11648/j.plant.20261401.11

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  • @article{10.11648/j.plant.20261401.11,
      author = {Zhen-Hao Feng},
      title = {Gathering Pearls from the Lethe: Nomenclatural Novelties for Nothotaxa},
      journal = {Plant},
      volume = {14},
      number = {1},
      pages = {1-23},
      doi = {10.11648/j.plant.20261401.11},
      url = {https://doi.org/10.11648/j.plant.20261401.11},
      eprint = {https://article.sciencepublishinggroup.com/pdf/10.11648.j.plant.20261401.11},
      abstract = {The integration of fine-scale taxonomic results into a comprehensive phylogenetic classification remains an ongoing challenge in modern systematics, particularly for infraspecific variants and interspecific hybrids, which are often overlooked. To address the resulting nomenclatural gaps and instabilities, this study provides a significant update by introducing numerous nomenclatural novelties for selected vascular plant genera, with a special focus on nothotaxa. In total, this work validates 452 nomenclatural novelties across 31 families, including Aizoaceae, Amaranthaceae, Amaryllidaceae, Apiaceae, Apocynaceae, Asteraceae, Berberidaceae, Cactaceae, Cistaceae, Crassulaceae, Cupressaceae, Cyperaceae, Equisetaceae, Fabaceae, Gesneriaceae, Melanthiaceae, Orchidaceae, Poaceae, Ranunculaceae and Saxifragaceae, ect. Among them are the establishment of 12 new nothogenera: × Osteolaena [Amaranthaceae], × Chrysania, × Cicuca, × Ozodes and × Psepaurea [Asteraceae], × Turdenicarpus [Cactaceae], × Astoxytropis [Fabaceae], × Eucongia [Gesneriaceae], × Pseudillium [Melanthiaceae], × Batepalum [Orchidaceae], × Rankwithia [Ranunculaceae] and × Mitarella [Saxifragaceae]; 59 nothoranks [five nothosubgenera, 42 nothosections, six nothosubsections and six nothoseries] under seven nothogenera [× Anacamptorchis, × Dactylocamptis, × Dactylodenia, × Gymnanacamptis, × Orchidactylorhiza, × Orchigymnadenia and × Orchimantoglossum] for the first time and as additions to five natural genera [Cattleya, Dactylorhiza, Paphiopedilum, Phalaenopsis and Pulsatilla], respectively; moreover, 360 new nomenclatural acts are proposed, covering 139 new combinations, 209 new combinaton and status, six new status, and six new replacement names; besides, nine names are synonymized under eight taxa. These changes reconcile hybrid taxonomy with contemporary phylogenetic insights, stabilize nomenclature and provide a crucial reference to support accurate identification, biodiversity documentation, and conservation efforts.},
     year = {2026}
    }
    

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  • TY  - JOUR
    T1  - Gathering Pearls from the Lethe: Nomenclatural Novelties for Nothotaxa
    AU  - Zhen-Hao Feng
    Y1  - 2026/01/27
    PY  - 2026
    N1  - https://doi.org/10.11648/j.plant.20261401.11
    DO  - 10.11648/j.plant.20261401.11
    T2  - Plant
    JF  - Plant
    JO  - Plant
    SP  - 1
    EP  - 23
    PB  - Science Publishing Group
    SN  - 2331-0677
    UR  - https://doi.org/10.11648/j.plant.20261401.11
    AB  - The integration of fine-scale taxonomic results into a comprehensive phylogenetic classification remains an ongoing challenge in modern systematics, particularly for infraspecific variants and interspecific hybrids, which are often overlooked. To address the resulting nomenclatural gaps and instabilities, this study provides a significant update by introducing numerous nomenclatural novelties for selected vascular plant genera, with a special focus on nothotaxa. In total, this work validates 452 nomenclatural novelties across 31 families, including Aizoaceae, Amaranthaceae, Amaryllidaceae, Apiaceae, Apocynaceae, Asteraceae, Berberidaceae, Cactaceae, Cistaceae, Crassulaceae, Cupressaceae, Cyperaceae, Equisetaceae, Fabaceae, Gesneriaceae, Melanthiaceae, Orchidaceae, Poaceae, Ranunculaceae and Saxifragaceae, ect. Among them are the establishment of 12 new nothogenera: × Osteolaena [Amaranthaceae], × Chrysania, × Cicuca, × Ozodes and × Psepaurea [Asteraceae], × Turdenicarpus [Cactaceae], × Astoxytropis [Fabaceae], × Eucongia [Gesneriaceae], × Pseudillium [Melanthiaceae], × Batepalum [Orchidaceae], × Rankwithia [Ranunculaceae] and × Mitarella [Saxifragaceae]; 59 nothoranks [five nothosubgenera, 42 nothosections, six nothosubsections and six nothoseries] under seven nothogenera [× Anacamptorchis, × Dactylocamptis, × Dactylodenia, × Gymnanacamptis, × Orchidactylorhiza, × Orchigymnadenia and × Orchimantoglossum] for the first time and as additions to five natural genera [Cattleya, Dactylorhiza, Paphiopedilum, Phalaenopsis and Pulsatilla], respectively; moreover, 360 new nomenclatural acts are proposed, covering 139 new combinations, 209 new combinaton and status, six new status, and six new replacement names; besides, nine names are synonymized under eight taxa. These changes reconcile hybrid taxonomy with contemporary phylogenetic insights, stabilize nomenclature and provide a crucial reference to support accurate identification, biodiversity documentation, and conservation efforts.
    VL  - 14
    IS  - 1
    ER  - 

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